| DEK58 |
is essential for |
endosperm development |
Zea mays |
| increased protein body size in line 107 |
is likely due to |
need to package substantial 19-kD α-zein in smaller number of protein bodies |
Zea mays |
| Gt1 promoter |
functions during the early stages of endosperm development but subsequently shows an increasing trend |
|
|
| type I MADS-box genes |
are regulated by |
MET1-dependent small interfering RNAs (p4-siRNAs) |
Arabidopsis thaliana |
| TtPARC6 expression |
is highest during |
early endosperm development |
Triticum turgidum |
| WT endosperm |
was |
nearly completely filled |
Zea mays |
| (ASG6, CRK2, AT1G70520) |
likely affects |
endosperm cell division and growth |
Zea mays |
| timing of endosperm initiation |
is critical for |
grain size |
|
| expression patterns of TtPDV1-1-A1 and TtPDV1-1-B1 |
differed from each other and from |
TtPARC6 expression patterns |
Triticum turgidum |
| cell number of the whole endosperm |
were significantly reduced in crk2 relative to |
WT |
Zea mays |
| (ASG6, CRK2, AT1G70520) endosperm cells |
were |
irregular |
Zea mays |
| promoter reporters of the MEGs (AGL36, AT5G26650) (AGL90, AT5G27960) and (AGL96, AT5G06500) |
displayed |
exclusively maternal up-regulation in (FIE2, FIS2, AT2G35670) and (EMB173, FIS1, MEA, SDG5, AT1G02580) mutants at 5 DAP |
Arabidopsis thaliana |
| transition to cellularization phase |
initiates |
cytokinesis |
|
| Endosperm Balance Number model |
could apply similar mechanism to |
imprinted PcG gene regulation of endosperm development |
|
| basal endosperm transfer layer (BETL) cells |
were undetectable in crk2 at 15 DAP, whereas present in WT |
more than two layers of BETL cells |
Zea mays |
| seeds derived from (TGS1, AT1G45231) FGs |
showed |
reduced proliferation in the endosperm |
Arabidopsis thaliana |
| expression of TtPDV1-2-A1 and TtPDV1-2-B1 |
mimics |
patterns of TtPARC6 expression |
Triticum turgidum |
| mutant plants (swn-3 /−; mea-3 /+) |
produced |
persistent coenocytic, noncellularized-type endosperm in seeds 4 and 5 DAP |
Arabidopsis thaliana |
| C2.1 genes (eight of 11) |
generally displayed higher activity in |
micropylar and peripheral regions of the coenocytic endosperm |
Arabidopsis thaliana |
| (AGL37, PHE1, AT1G65330) promoter reporter lines |
detected |
similar up-regulation pattern |
Arabidopsis thaliana |
| moderate mis-expression of the target genes |
would result in |
moderate endosperm abnormality or phenotype |
Arabidopsis thaliana |
| B-type granules |
are initiated later during |
endosperm development |
Triticum aestivum |
| TtARC6 expression in the endosperm |
is about tenfold higher than |
TtPARC6 expression |
Triticum turgidum |
| TtPDV1-1-A1 and TtPDV2-A1 expression patterns |
had similar |
patterns to TtARC6 |
Triticum turgidum |
| expression levels of BETL-specific marker genes |
were significantly reduced in crk2 mutant relative to |
WT |
Zea mays |
| interploidy and interspecific crosses |
show |
imbalanced parental genome dosages |
Arabidopsis thaliana |
| C1 genes |
showed lower mRNA levels during |
coenocytic endosperm development (1–3 DAP) |
Arabidopsis thaliana |
| C2 genes |
were up-regulated in |
mea-3;swn-3 mutant corresponding to the period of endosperm cellularization |
Arabidopsis thaliana |
| (FIE2, FIS2, AT2G35670) mutant |
produces phenotype similar to |
cellularization timing defects in species hybrids |
Arabidopsis thaliana |
| increased 27-kD γ-zein in QPM |
is consistent with suggested role in |
protein body formation in QPM |
Zea mays |
| coenocytic program |
is repressed at |
onset of cellularization |
Arabidopsis thaliana |
| FERTILIZATION-INDEPENDENT SEED (FIS)-Polycomb Repressive Complex2 (PRC2) |
is composed primarily of |
(FIE2, FIS2, AT2G35670) MEDEA (EMB173, FIS1, MEA, SDG5, AT1G02580) FERTILIZATION-INDEPENDENT ENDOSPERM (FIE, FIE1, FIS3, AT3G20740) and Arabidopsis homolog of MULTICOPY SUPPRESSOR OF IRA1 (ATMSI1, MEE70, MSI1, AT5G58230) |
Arabidopsis thaliana |
| (AGL36, AT5G26650) and (AGL37, PHE1, AT1G65330) promoter reporter genes |
showed |
maternally restricted activity at 2 DAP |
Arabidopsis thaliana |
| K0326Y Quality Protein Maize (QPM) deletion mutant |
abolished |
vitreous endosperm formation |
Zea mays |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) |
is |
endosperm-expressed gene with locus-specific imprinting |
Arabidopsis thaliana |
| (AGL62, AT5G60440) |
suppresses |
cellularization |
Arabidopsis thaliana |
| AGAMOUS-LIKE62 (AGL62, AT5G60440) |
promotes |
coenocytic development |
Arabidopsis thaliana |
| (AGL90, AT5G27960) mutant |
alleviates |
endosperm-mediated postzygotic barriers between Arabidopsis and Arabidopsis arenosa |
Arabidopsis thaliana; Arabidopsis arenosa |
| K0326Y Quality Protein Maize (QPM) deletion mutant |
has protein body size similar to |
wild type |
Zea mays |
| coenocytic endosperm |
differentiates into |
micropylar domain |
Arabidopsis thaliana |
| C2.1 genes (seven of 11) |
were rapidly down-regulated in |
all three endosperm compartments |
Arabidopsis thaliana |
| 16-kD γ-zein |
is not sufficient for supporting |
normal level of protein body initiation |
Zea mays |
| irregular undulating protein body shape |
may be due to |
improper packaging of hydrophobic 19-kD α-zein |
Zea mays |
| (AGL23, AT1G65360) |
showed |
more delayed down-regulation pattern |
Arabidopsis thaliana |
| all five C2.2 genes |
displayed reduced activity in the micropylar and peripheral endosperm but maintained high activity in |
chalazal endosperm |
Arabidopsis thaliana |
| Ploidy barrier to endosperm hypothesis |
was proposed to explain |
endosperm incompatibility in maize |
Zea mays |
| timing of transition outside permissible range |
causes |
seed abortion |
|
| γ-zein deletion |
affects |
protein body initiation, morphology, and zein distribution |
Zea mays |
| global DNA hypomethylation |
causes |
abbreviated coenocytic phase |
Arabidopsis thaliana |
| all five C2.2 genes |
typically displayed higher GFP activity in |
both the micropylar and chalazal regions or were expressed predominantly in the chalazal region |
Arabidopsis thaliana |
| paternal silencing of C2 MEGs |
can be maintained exclusively through |
5′-flanking region of the gene by a FIS-PRC2-independent mechanism |
Arabidopsis thaliana |
| type I MADS-box gene family |
is partly required for |
regulating coenocytic development |
Arabidopsis thaliana |
| full-length gene fusions for (AGL36, AT5G26650) and (AGL37, PHE1, AT1G65330) |
displayed |
expected patterns of expression |
Arabidopsis thaliana |
| expression from single allele of 27- and 50-kD γ-zein genes |
is haploinsufficient for |
full o2 endosperm modification |
Zea mays |
| hypomethylation of paternal genome |
causes |
abbreviated coenocytic phase |
Arabidopsis thaliana |
| FIS-PRC2 complex |
may dissociate from |
C2 loci during coenocytic endosperm phase |
|
| T-DNA insertion lines in rice homologs of three PcG genes |
show |
no autonomous endosperm development |
Oryza sativa |
| AGL62–GFP signals |
do not disappear prior to |
cellularization in PcG mutants |
Arabidopsis thaliana |
| up-regulated pattern of C2 genes |
corresponded to |
autonomous (3 DAE) and noncellularized endosperm (4–5 DAP) phenotypes |
Arabidopsis thaliana |
| both the paternal and maternal alleles of most of the biallelically expressed C2 genes |
were up-regulated at 5 DAP in |
mutants when tested as promoter reporters |
Arabidopsis thaliana |
| line 107 protein bodies |
have irregular, undulating surfaces |
protein body morphology |
Zea mays |
| complete absence of 27-kD γ-zein |
results in even more severe suppression of |
protein body initiation |
Zea mays |
| pheres1 ( (AGL37, PHE1, AT1G65330) also known as ) mutant |
alleviates |
endosperm-mediated postzygotic barriers between Arabidopsis and Arabidopsis arenosa |
Arabidopsis thaliana; Arabidopsis arenosa |
| regulation of cellularization |
has functional link with |
genomic imprinting |
|
| PHERES1 (AGL37, PHE1, AT1G65330) |
is suggested to regulate |
endosperm development |
Arabidopsis thaliana |
| C2.1 and C2.2 genes |
are programmed for |
differential expression in distinct domains of coenocytic endosperm prior to cellularization |
Arabidopsis thaliana |
| (AGL37, PHE1, AT1G65330) |
is |
endosperm-expressed gene with locus-specific imprinting |
Arabidopsis thaliana |
| endosperm development |
must be regulated by |
imprinted Polycomb group (PcG) genes through type-I MADS-box proteins |
Arabidopsis thaliana |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) (EZA1, SDG10, SWN, AT4G02020) double mutant |
produces |
persistent coenocytic endosperm |
|
| Endosperm Balance Number hypothesis |
was proposed to explain |
endosperm incompatibility in potatoes |
Solanum tuberosum |
| timing of transition outside permissible range |
causes |
endosperm abortion |
|
| FIS-PRC2 complex |
maintain |
locus-specific, paternal or maternal imprinting of some endosperm-expressed genes |
Arabidopsis thaliana |
| AGAMOUS-LIKE62 (AGL62, AT5G60440) |
regulates |
coenocytic endosperm development |
Arabidopsis thaliana |
| PHERES1 (AGL37, PHE1, AT1G65330) antisense transgenic plants |
show |
no obvious phenotype in endosperm development |
Arabidopsis thaliana |
| type I MADS-box genes |
are up-regulated in |
crosses with excess paternal genome |
Arabidopsis thaliana |
| mRNA of the GluA-2 gene |
peaked at approximately |
10 DAF |
|
| mutant plants (swn-3 /−; mea-3 /+) |
produced |
autonomous endosperm at 3 DAE |
Arabidopsis thaliana |
| C2.1 and C2.2 genes |
exhibited |
differential mRNA accumulation in the endosperm compartments |
Arabidopsis thaliana |
| osfie1 mutant |
did not show |
autonomous endosperm |
Oryza sativa |
| RNAi suppression of 27-kD γ-zein synthesis |
resulted in suppression of |
protein body initiation |
Zea mays |
| type I MADS-box genes |
are regulated by |
FIS-PRC2-mediated histone modification |
Arabidopsis thaliana |
| FIS-PRC2 |
required to repress expression of C2 genes before |
endosperm initiation |
|
| endosperm nuclei after fertilization |
undergo many rounds of |
mitosis without cytokinesis |
|
| (AGL62, AT5G60440) |
is involved in regulation of |
endosperm cellularization |
Arabidopsis thaliana |
| coenocytic program |
is repressed prior to |
fertilization |
Arabidopsis thaliana |
| species II as tetraploid plant in crossing combination of species I and species II |
results in dosage of paternally derived genome being |
twice in the endosperm |
|
| species I as tetraploid in crossing combination with species II |
results in dosage of maternally supplied chromosomes being |
doubled |
|
| dek58 mutant kernel at 9 DAP |
develops |
small endosperm |
Zea mays |
| coenocytic endosperm |
differentiates into |
chalazal domain |
Arabidopsis thaliana |
| (EZA1, SDG10, SWN, AT4G02020) and (EMB173, FIS1, MEA, SDG5, AT1G02580) FIS-PRC2 complexes |
perform partially redundant roles in regulating |
initiation of endosperm development upon fertilization |
Arabidopsis thaliana |
| protein body number in line 107 |
is dramatically reduced compared with |
wild type and QPM |
Zea mays |
| type I MADS-box single-gene mutants |
have |
no discernible phenotype in the endosperm |
Arabidopsis thaliana |
| imprinted components in complex |
renders complex sensitive to |
imbalanced parental genome dosage |
|
| imprinted Polycomb group genes (PcG) |
affects timing of |
cellularization during endosperm development |
Arabidopsis thaliana |
| 12 DAP stage in maize endosperm |
is associated with |
cell division |
Zea mays |
| different combinations of epigenetic marks |
determine |
activity status of maternal alleles in developing endosperm |
Arabidopsis thaliana |
| number of differentially expressed genes (DEGs) |
gradually decreased |
endosperm development progression |
Triticum urartu |
| mn1 mutant |
showed |
reduced cell number during endosperm development |
Zea mays |
| expression profile of three TaNAC77 homoeologs |
started after |
5 (DPA, AT5G02470) |
Triticum aestivum |
| maize endosperm |
was sampled during |
multiple developmental stages |
Zea mays |
| Zma-miR159k-3p |
is highly and endogenously expressed in |
developing maize endosperm |
Zea mays |
| T-DNA insertion mutants of OsFIE1 |
shows |
no autonomous endosperm development |
Oryza sativa |
| finding |
was consistent with |
methylation cluster analysis, which showed that the end of storage compound accumulation in the endosperm clustered close to the 3–5 DAP time point |
Zea mays |
| endosperm-expressed genes |
likely undergo strong transcriptional regulation during |
7 to 10 DAP period |
Zea mays |
| miRNA-mediated post-transcriptional regulation |
occurs during |
maize endosperm development |
Zea mays |
| dark-brown substance in HFL1 caryopsis |
was observed mainly in |
later stage of endosperm development (more than 25 DAF) |
Oryza sativa |
| endosperm sensitivity to parental genome dosage |
suggests |
some regulators of endosperm development are subjected to imprinting |
Arabidopsis thaliana |
| (AGL62, AT5G60440) mutant |
alleviates |
endosperm-mediated postzygotic barriers between Arabidopsis and Arabidopsis arenosa |
Arabidopsis thaliana; Arabidopsis arenosa |
| type I MADS-box genes |
are expressed in |
coenocytic endosperm |
Arabidopsis thaliana |
| genetic networks |
regulate |
endosperm initiation |
Oryza sativa |
| genetic and epigenetic variations in maternal or paternal contributions |
result in |
reciprocal effects in endosperm development |
|
| mitosis coupled with cell division |
is |
one of three cell cycle types during endosperm development |
Oryza sativa L. |
| syncytium endosperm of nuf2-1/+ and nuf2-3/+ mutant ovules |
failed to undergo |
normal cellularization |
Arabidopsis thaliana |
| number of expressed genes and percentage of genes with medium, high and very high expression levels at 25 (DPA, AT5G02470) |
were higher than |
number of expressed genes and percentage of genes with medium, high and very high expression levels at 20 (DPA, AT5G02470) |
Triticum urartu |
| parental conflict |
mediates |
epigenetic molecular machinery in endosperm |
|
| endosperm of (CYL1, NAGLU, AT5G13690) seeds |
was composed of |
12–20 syncytial nuclei |
|
| syncytium endosperm of nuf2-1/+ and nuf2-3/+ mutant ovules |
was composed of |
free nuclei of different sizes |
Arabidopsis thaliana |
| paternal excess endosperm (2m:2p) |
leads to |
nuclei over-proliferation |
|
| AroDH-2 |
is playing little or no functional role in |
the endosperm |
Zea mays |
| early phase of endosperm development |
is |
3–5 DAP |
Zea mays |
| distinct cluster of 20–25 DAP |
was consistent with |
period of endosperm maturation and cell death |
Zea mays |
| syncytium formation |
is followed by |
cellularization |
Zea mays |
| transcriptome changes |
may activate |
starch accumulation |
Zea mays |
| maternal alleles marked by triple repressive marks H3K27me3/H3K9me2/CHGm |
tend to remain silenced during |
endosperm development |
Arabidopsis thaliana |
| genetic networks |
regulate |
interactions between embryo and endosperm |
Oryza sativa |
| seven gene clusters (C1–C7) |
have |
different expression trends during endosperm development |
Triticum urartu |
| starch accumulation |
begins around |
6–9 DAP |
Zea mays |
| cellularization |
precedes starch grain accumulation in |
maize endosperm |
Zea mays |
| epigenetic regulators |
have key function in |
parental control of endosperm development |
|
| Molecular machinery and regulatory architecture governing endosperm development |
is proposed to have evolved under |
parental conflict |
|
| srn phenotype |
might be caused by |
onset of autonomous development of the central cell |
|
| Orysa; (ICK6, KRP3, AT5G48820) mRNA |
was barely detectable in |
the primary cellularized cells at 3 DAF |
Oryza sativa |
| developmental stages of 7–15 DAP |
are |
periods during which the major cell types are formed and storage materials are rapidly accumulated |
Zea mays |
| Orysa; (ICK6, KRP3, AT5G48820) expression |
is down-regulated after |
cellularization |
Oryza sativa L. |
| syl |
is not associated to |
maternal effect |
|
| 3.5 DAE |
allows observation of |
autonomous endosperm development |
|
| mitosis without cytokinesis |
results in |
syncytium formation |
Oryza sativa L. |
| overexpression or knockdown of Orysa; (ICK6, KRP3, AT5G48820) |
can be used to engineer |
rice plants with enhanced or reduced levels of Orysa; (ICK6, KRP3, AT5G48820) expression |
Oryza sativa |
| delayed fertilization of emasculated wild-type ovules |
resulted in increased frequency of |
srn phenotype |
|
| early shrinkage of endosperm tissue |
initiates at |
late heart stage |
Arabidopsis thaliana |
| (NF-YC12, AT5G38140) |
directly binds to the promoter of |
FLOURY ENDOSPERM6 |
Oryza sativa |
| positive signal from fertilization of egg cell |
might promote |
onset of endosperm cell development |
Arabidopsis thaliana |
| rice seeds at 2 DAF |
contain |
numerous syncytial nuclei in the periphery of the central cell |
Oryza sativa |
| Orysa; (ICK6, KRP3, AT5G48820) |
determines |
final seed size |
Oryza sativa |
| controlling cell cycle in syncytial endosperm by manipulation of Orysa; (ICK6, KRP3, AT5G48820) |
could offer the potential to improve |
seed size |
Oryza sativa |
| diameter of TIP3-GFP-labelled PSVs |
fell to ~4.92 μm at |
12 DAP |
Hordeum vulgare |
| partial enclosure of protein bodies by TIP3-GFP-labelled membranes |
suggested |
membrane integrity in starchy endosperm may be affected during maturation |
Hordeum vulgare |
| genetic and epigenetic controls |
play crucial roles in |
rice endosperm development |
Oryza sativa |
| syl mutant |
exhibits |
srn phenotype |
|
| syncitial ring of nuclei |
surrounds |
central vacuole |
|
| vacuolation of protein storage vacuoles |
takes place later in |
non-micropylar endosperm and embryo |
Arabidopsis thaliana |
| cell wall composition in Brachypodium central endosperm and nucellar epidermis |
is more closely related to |
cell wall composition in barley and oats |
Brachypodium distachyon; Hordeum vulgare; Avena sativa |
| endosperm development in cereals |
is |
distinctive process |
|
| vacuolar compartments in subaleurone |
became smaller by |
10 DAP |
Hordeum vulgare |
| autonomous endosperm development in absence of fertilization |
is associated with |
FERTILIZATION INDEPENDENT SEED (FIS) mutant class |
Arabidopsis thaliana |
| syl mutant ovules |
exhibit |
autonomous development |
|
| syncytial endosperm |
contains |
16 small nuclei |
|
| rice seeds at 3 DAF |
show |
two layers of walled cells at the periphery of the endosperm |
Oryza sativa |
| Arath; (ACK2, ICK7, KRP4, AT2G32710) |
showed |
endosperm-preferred expression |
Arabidopsis thaliana |
| timing of cellularization |
correlates with |
extent of nuclear proliferation |
Oryza sativa L. |
| mitosis coupled with cell division and endoreduplication |
have been characterized during |
later stages of endosperm development |
Oryza sativa L. |
| genomic imprinting |
plays key role in |
endosperm development |
|
| absence of srn-dependent signaling |
would cause |
initiation of central cell development |
|
| a series of divisions without cytokinesis |
produces |
multinucleate cell known as a syncytium |
Oryza sativa |
| expression of Arath; (ACK2, ICK7, KRP4, AT2G32710) |
is not limited in |
endosperm |
Arabidopsis thaliana |
| (FIE2, FIS2, AT2G35670) transcripts |
were affected by |
(ATCSP4, ATGRP2B, GRP2B, AT2G21060) overexpression |
Arabidopsis thaliana |
| endosperm development |
progresses via |
division of central cell-derived triploid nucleus |
|
| disruption of asters |
impairs |
endosperm proliferation |
|
| functionally distinct subdomains |
include |
central starchy endosperm |
|
| multinucleate endosperm stage |
is associated with |
endosperm proliferation and cell differentiation |
Triticum aestivum |
| genetic networks |
regulate |
aleurone layer specification |
Oryza sativa |
| differential screening, mutant screening, microdissection, and microarray experiments |
identified |
genes preferentially expressed in syncytial endosperm |
Arabidopsis thaliana; Hordeum vulgare |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) mRNA |
is highly accumulated during early stage of embryogenesis in |
35S:NTAP:AtCSP4-3 plants |
Arabidopsis thaliana |
| (CYL1, NAGLU, AT5G13690) mutant endosperm |
contains |
up to 20 nuclei |
|
| 26 285 genes |
were expressed in |
at least one of the five time points of endosperm development |
Triticum urartu |
| paternal excess endosperm (2m:2p) |
leads to |
failure to cellularize |
|
| diameter of TIP3-GFP-labelled PSVs |
rose to a maximum of 77.2 μm at |
8 DAP |
Hordeum vulgare |
| size of TIP3-GFP-labelled vacuolar compartments |
was reduced by ~33% between |
8 and 10 DAP |
Hordeum vulgare |
| protein storage vacuoles (PSVs) |
fate is little known in |
developing cereal endosperm |
|
| imprinted genes strongly associated with (ATNACK2, NACK2, TES, AT3G43210) |
have broad expression across |
endosperm development and domains |
|
| mode of development |
was reminiscent of |
autonomous endosperm development observed in fis -class mutants |
|
| proliferating tissue from the emasculated ovule |
expressed |
endosperm marker (FIE2, FIS2, AT2G35670) |
|
| delayed pollination for 1.5 DAE |
coupled with |
autonomous development |
|
| regions that lack aleurone in (ACR4, CR4, AT3G59420) mutant |
have peripheral part consisting of |
starchy endosperm cells |
Zea mays |
| des5 mutant phenotype |
exhibits |
morphologically defective starchy endosperm cells |
Hordeum vulgare |
| aleurone in Brachypodium |
is not regionally differentiated as in |
aleurone in wheat |
Brachypodium distachyon; Triticum aestivum |
| experimental set-up including isolated embryo sacs on a medium favourable for in vitro pollen tube germination |
allowed |
recording of the first series of syncytial nuclei division in the endosperm |
Torenia fournieri |
| TIP3-GFP-labelled PSVs |
remained morphologically stable in |
aleurone |
Hordeum vulgare |
| protein bodies in the maturing starchy endosperm |
no longer fully surrounded by |
TIP3-GFP-labelled membranes |
Hordeum vulgare |
| (AGL62, AT5G60440) mutant seeds |
exhibit |
early syncytial endosperm overproliferation |
Arabidopsis thaliana |
| anticlinal divisions in the periphery |
accommodate |
enlargement of the developing grain |
|
| bareback and naked mutants |
have |
opaque and often white endosperm with floury texture |
Zea mays |
| Mutator-tagged opaque 140 (mto140) |
is |
opaque endosperm mutant |
Zea mays |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) transcripts |
were affected by |
(ATCSP4, ATGRP2B, GRP2B, AT2G21060) overexpression |
Arabidopsis thaliana |
| loss-of-function mutant of the FIS gene |
perturbs |
endosperm development |
Arabidopsis thaliana |
| BR-dependent promotion of seed growth |
impacts |
early development of the coenocytic endosperm |
|
| imprinted genes with little or no TE association |
have restricted expression in |
specific stages or regions of the endosperm |
|
| seeds with viable paternal excess (containing 2m:2p endosperms) |
are larger than |
seeds from balanced crosses (2m:1p) |
Arabidopsis thaliana |
| differences in Brachypodium endosperm development |
may reflect |
phylogenetic position between Triticeae and rice |
Brachypodium distachyon |
| cereal endosperm aleurone layers |
rely on |
positional information |
|
| fate of TIP3-GFP-labelled PSVs |
was dependent on |
cell layer |
Hordeum vulgare |
| rupture of small vacuoles |
caused |
release of TIP3-GFP-labelled vesicles and membranes |
Hordeum vulgare |
| internal membranes in barley endosperm protein storage vacuoles (PSVs) |
revealed by labelling of |
membrane components |
Hordeum vulgare |
| (ATDPB2, CYL2, DPB2, AT5G22110) mutant endosperm |
contains |
1–4 nuclei |
|
| TIP3-GFP signal |
progressively weaker in |
subaleurone and starchy endosperm |
Hordeum vulgare |
| endosperm development |
is initiated by |
series of synchronous cell cycles in absence of cytokinesis |
Arabidopsis thaliana |
| H3.3-mRFP1 signal |
persists until |
third series of mitoses |
Arabidopsis thaliana |
| des5 mutant seeds |
have |
loose and shrunken endosperm |
Hordeum vulgare |
| mto140 mature kernel |
manifests |
marked reduction in the thickness of the vitreous endosperm layer |
Zea mays |
| endosperm of (CYL1, NAGLU, AT5G13690) seeds |
is comparable to |
wild-type endosperm at the one cell stage |
|
| pollination at 3.5 DAE |
resulted in decreased |
srn phenotype percentage |
|
| endosperm development |
includes phase of |
cellularization |
Oryza sativa L. |
| ABA |
inhibits |
vacuolation |
Arabidopsis thaliana |
| duration of the syncytial phase |
correlates with |
extent of nuclear proliferation |
Oryza sativa L. |
| early stages of maize endosperm development (9–12 days after pollination) |
no protein complexes were detected in |
either genotype |
Zea mays |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) and (FIE2, FIS2, AT2G35670) expression |
was affected during |
early seed development stages |
Arabidopsis thaliana |
| one A-type granule |
is initiated early during |
endosperm development |
Triticum aestivum |
| altered A-type granule morphology |
manifests early during |
grain development |
Triticum turgidum |
| endosperm development in (ASG6, CRK2, AT1G70520) |
was |
delayed |
Zea mays |
| average size of endosperm cells in (ASG6, CRK2, AT1G70520) |
was approximately |
40% of that in WT |
Zea mays |
| mto140 |
is similar to |
opaque5 (o5) and opaque9 (o9) mutants |
Zea mays |
| differences in cell morphology and protein storage vacuole (PSV) properties |
likely reflecting |
centripetal increase of cell age |
Hordeum vulgare |
| protein complexes |
once formed, are present in the amyloplast until |
relatively late in development (up to 35 days after pollination) |
Zea mays |
| 4n self-pollinated seeds |
show |
characteristic nuclear/cytoplasmic domains of syncytial-stage endosperm until 2 DAP |
Oryza sativa |
| syncytial endosperm development |
has been suggested to have evolved independently in |
monocots and eudicots |
|
| global DNA hypomethylation in the endosperm |
is established by active demethylation by demeter (DME) |
active demethylation by demeter (DME, EMB1649, AT5G04560) |
Arabidopsis thaliana |
| (DDM2, DMT01, DMT1, MET1, MET2, METI, AT5G49160) |
interacts directly with |
(EMB173, FIS1, MEA, SDG5, AT1G02580) |
Arabidopsis thaliana |
| histone methylation introduced by the FIS-PRC2 |
is sufficient to repress |
genes important for the initiation of endosperm development |
Arabidopsis thaliana |
| small protein bodies |
fused into |
larger aggregates |
Hordeum vulgare |
| protein storage vacuoles (PSVs) in the subaleurone |
undergo fusion followed by |
formation of larger protein bodies |
Hordeum vulgare |
| reciprocal cross between female diploid Arabidopsis thaliana (2n) and male tetraploid Arabidopsis thaliana (4n) with paternal genome excess |
shows |
delayed cellularization |
Arabidopsis thaliana |
| OsMADS87 gene |
is over-expressed in |
2n × 4n cross at 5 DAP |
Oryza sativa |
| cessation of rapid nuclear proliferation in 2n × 4n cross between 1.5 and 2.0 DAP |
is probably because of |
endosperm abnormality |
Oryza sativa |
| underlying molecular mechanisms |
may not be conserved between |
Arabidopsis thaliana and Oryza sativa |
Arabidopsis thaliana; Oryza sativa |
| TIP3-GFP-positive membrane structures |
movement was followed in |
time-course experiment |
Hordeum vulgare |
| cereal endosperm cells |
have been characterized by |
histochemical fluorescence and electron microscopy studies |
|
| TIP3-GFP-labelled protein storage vacuoles (PSVs) in the central starchy endosperm |
reduced in size considerably by |
10 days after pollination (10 DAP) |
Hordeum vulgare |
| cellularized structure at the anterior pole |
should be |
diploid (n = 10) |
Arabidopsis thaliana |
| ten centromeres of maternal chromosomes |
remained unmarked by |
paternal HTR12-GFP during first three syncytial divisions |
Arabidopsis thaliana |
| TIP3-GFP-labelled PSVs |
became smaller and less regular in shape by |
12 DAP |
Hordeum vulgare |
| TIP3-labelled protein storage vacuoles (PSVs) of the subaleurone at 10 days after pollination (10 DAP) |
characterized by |
dynamic reshaping events |
Hordeum vulgare |
| starch granules |
become more prominent during |
maturation |
Hordeum vulgare |
| central starchy endosperm of barley beyond 12 DAP |
contained |
small TIP3-GFP-labelled PSVs |
Hordeum vulgare |
| radial microtubule systems (RMSs) |
organize |
nuclear cytoplasmic domains (NCDs) |
|
| small cellular structure |
was eventually considered as |
cellularized derivative of endosperm |
Arabidopsis thaliana |
| TIP3-GFP signal |
was already localized to |
vacuolar membranes in earlier undifferentiated endosperm |
Hordeum vulgare |
| two small vacuoles surrounding protein bodies |
fused to form |
larger TIP3-GFP-labelled PSV |
Hordeum vulgare |
| observations of fusion and collapse processes |
were likely to contribute to |
morphological changes of TIP3-GFP-labelled PSVs during endosperm development |
Hordeum vulgare |
| collapsing subaleurone protein storage vacuoles (PSVs) at the same developmental stage |
considered together with |
protein bodies in central starchy endosperm lacking complete membranes |
Hordeum vulgare |
| six of these seven AGLs |
are |
downregulated at the transition from syncytial to cellular endosperm growth |
Arabidopsis thaliana |
| endosperm nuclei |
increase until |
bent-cotyledon stage |
Arabidopsis thaliana |
| ER-derived membranes |
may be associated with |
individual and clustered protein bodies within subaleurone protein storage vacuoles (PSVs) |
Hordeum vulgare |
| dynamic reshaping events in subaleurone protein storage vacuoles (PSVs) |
not so profound in |
other layers |
Hordeum vulgare |
| absence of pronounced ER network |
indicated |
alterations of ER membrane system |
Hordeum vulgare |
| starch granules in the starchy endosperm |
become more prominent in |
starchy endosperm |
Hordeum vulgare |
| endosperm in interploidy crosses |
shows |
smaller maximum cross-sectional area at early stage of seed development |
Oryza sativa |
| nuclear proliferation in the syncytium stage |
is |
one of two separable components of the post-zygotic hybridization barrier |
Oryza sativa |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) |
is involved in the repression of central cell proliferation and endosperm development in the absence of fertilization |
central cell proliferation and endosperm development |
Arabidopsis thaliana |
| at least partially functional repressive FIS-PRC2 |
is formed in the absence of |
(DDM2, DMT01, DMT1, MET1, MET2, METI, AT5G49160) |
Arabidopsis thaliana |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) (EZA1, SDG10, SWN, AT4G02020) double mutants |
show |
incomplete penetrance of fis phenotype |
Arabidopsis thaliana |
| knockout of AtRFC4 |
results in |
division of syncytial endosperm free nuclei |
Arabidopsis thaliana |
| unusually prominent cell walls in Brachypodium central endosperm and nucellar epidermis |
may act as |
storage material |
Brachypodium distachyon |
| syncitial ring of nuclei |
divide and cellularize in |
concerted sequence of anticlinal and periclinal cell divisions |
|
| diameter of TIP3-GFP-labelled PSVs |
fell to ~7.97 μm at |
10 DAP |
Hordeum vulgare |
| protein bodies in the starchy endosperm during later development |
appeared only partially enclosed by |
TIP3-GFP-labelled membranes |
Hordeum vulgare |
| fertilized wt endosperm |
is |
triploid (3N = 15) |
Arabidopsis thaliana |
| TIP3-GFP-labelled protein storage vacuoles (PSVs) in the subaleurone and central starchy endosperm |
reduced in size during |
development |
Hordeum vulgare |
| C2 genes |
represent |
endosperm-specific subclade of type I MADS-box genes |
Arabidopsis thaliana |
| (AGL96, AT5G06500) promoter reporter genes |
showed |
maternally restricted activity at 2 DAP |
Arabidopsis thaliana |
| PSVs from nearby aleurone layers |
remained unaffected in |
all cases |
Hordeum vulgare |
| chalazal compartment |
cellularizes late as compared with |
other endosperm compartments |
Arabidopsis thaliana |
| ABNORMAL LEAF-SHAPE1 (ALE, ALE1, AT1G62340) |
regulates |
degeneration of micropylar endosperm |
|
| syncytium |
undergoes |
cellularization and maturation |
Arabidopsis thaliana |
| (AGL62, AT5G60440) suppression of cellularization |
results in prolonged expression of the target genes of the PcG complex potentially delaying |
endosperm cellularization |
Arabidopsis thaliana |
| barley endosperm |
is fully cellularized by |
8 days after pollination |
Hordeum vulgare |
| 27-kD γ-zein |
has suggested role in |
endosperm modification in QPM |
Zea mays |
| swn-3 /−; mea-3 /+ double-mutant plants |
produce |
enhanced autonomous endosperm phenotype |
Arabidopsis thaliana |
| maternal and paternal alleles of the PEG (AGL37, PHE1, AT1G65330) full-length gene fusion |
were up-regulated at 5 DAP in |
(FIE2, FIS2, AT2G35670) and (EMB173, FIS1, MEA, SDG5, AT1G02580) mutant seeds |
Arabidopsis thaliana |
| FERTILIZATION INDEPENDENT SEED2 (FIE2, FIS2, AT2G35670) |
is |
component of PRC2 Polycomb protein complex |
Arabidopsis thaliana |
| starch granule size |
increased to ~11.45 μm at |
10 DAP |
Hordeum vulgare |
| TIP3-GFP-labelled PSVs in central starchy endosperm |
also reduced in size |
during development |
Hordeum vulgare |
| paternal MEDEA (EMB173, FIS1, MEA, SDG5, AT1G02580) -GUS expression in endosperm |
increased progressively |
during successive rounds of nuclei divisions |
Arabidopsis thaliana |
| C2 genes |
were mostly down-regulated as |
endosperm became cellularized |
Arabidopsis thaliana |
| MULTICOPY SUPPRESSOR OF IRA1 (ATMSI1, MEE70, MSI1, AT5G58230) |
is |
component of PRC2 Polycomb protein complex |
Arabidopsis thaliana |
| HvCSLC1 and HvCSLC2 transcript levels |
become undetectable by |
6 days after pollination |
Hordeum vulgare |
| rrp4-1 mutant endosperm |
developed to varying degrees but never past |
cellularization stage |
Arabidopsis thaliana |
| nucellus degeneration in interspecific cross 2x O. sativa × 2x O. longistaminata |
occurred precociously by |
2 DAP |
Oryza sativa; Oryza longistaminata |
| endosperm of O. sativa × O. longistaminata cross |
showed normal |
developmental transitions including timing of cellularization |
Oryza sativa; Oryza longistaminata |
| hypomethylation of maternal genome |
causes |
prolonged coenocytic phase |
Arabidopsis thaliana |
| FIS-PRC2 complex |
regulates |
type I MADS-box genes |
Arabidopsis thaliana |
| FIS-PRC2 |
expressed in |
coenocytic endosperm |
|
| imprinting |
suggested as underlying cause of |
abnormal endosperm development in crosses with imbalanced parental genome dosage |
|
| FIS-PRC2-mediated repression of C2 subclade |
is necessary to restrict |
C2 gene expression to the coenocytic phase of endosperm development |
Arabidopsis thaliana |
| FIS-PRC2 |
required to repress expression of C2 genes during |
endosperm cellularization |
|
| timing of phase transition in endosperm |
is important for |
seed development |
|
| effects on timing of cellularization in interspecific and interploidy crosses in rice |
are similar to |
effects found previously in interspecific crosses in rice |
Oryza sativa |
| interploidy crosses |
are consistent with |
results reported for interploidy and interspecies crosses in Arabidopsis thaliana |
Oryza sativa; Arabidopsis thaliana |
| Mutator insertion in zmAroDH-3 |
is associated with |
opaque endosperm phenotype |
Zea mays |
| embryo sheath in pTE2:KIR1 lines |
appears unaffected |
in pTE2:KIR1 lines |
Arabidopsis thaliana |
| Emb12 |
is not essential for |
endosperm development |
Zea mays |
| endosperm free nuclei in mutants at one-celled embryo proper stage |
have no more than |
eight free nuclei |
Arabidopsis thaliana |
| PHERES1 (AGL37, PHE1, AT1G65330) |
deregulation of |
endosperm development |
Arabidopsis thaliana; Arabidopsis arenosa |
| endosperm tissue in interploidy cross 4x O. sativa × 2x O. sativa |
was very |
small |
Oryza sativa |
| post-hybridization barrier |
has two major components |
altered timing of cellularization |
Arabidopsis thaliana |
| endosperm in Arabidopsis thaliana |
supports |
embryonic growth |
Arabidopsis thaliana |
| loss of Emb12 function |
has minimal impact on |
endosperm starch, lipid and storage protein accumulation |
Zea mays |
| (DDM2, DMT01, DMT1, MET1, MET2, METI, AT5G49160) mutant and mutants in various FIS-PRC2 components |
show different and partially overlapping effects on de-repression of |
genes targeted by both histone and DNA methylation |
Arabidopsis thaliana |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) gene |
is involved in |
endosperm development |
Arabidopsis thaliana |
| living central endosperm and aleurone cells |
have |
regularly shaped round nuclei |
Arabidopsis thaliana |
| OsMADS87 expression patterns in different crosses |
are consistent with |
patterns of delayed or precocious endosperm cellularization |
Oryza sativa; Oryza longistaminata |
| seed abortion phenotypes |
result from |
bidirectional alterations of endosperm development |
Oryza sativa; Oryza longistaminata |
| cellular endosperm |
is |
endosperm type distinct from nuclear-type endosperm |
Solanum section Lycopersicon |
| MEDEA (EMB173, FIS1, MEA, SDG5, AT1G02580) Pc-G complex |
controls |
endosperm growth and proliferation |
Arabidopsis thaliana |
| paternal HTR12-GFP |
is not removed from |
endosperm nuclei during first three syncytial divisions |
Arabidopsis thaliana |
| presence of intravacuolar TIP3-GFP-positive membranes |
indicated that |
reshaping events must have taken place |
Hordeum vulgare |
| (ATMSI1, MEE70, MSI1, AT5G58230) autonomous endosperm |
is |
diploid |
Arabidopsis thaliana |
| (AGL62, AT5G60440) mutant seeds |
results in |
death |
Arabidopsis thaliana |
| suppression of the AGL cluster at 5 DAP |
is critical for |
successful transition from the syncytial to cellularized stages of seed development |
Arabidopsis thaliana; Arabidopsis arenosa |
| DAPI-stained nuclei in zou-4 mutant endosperm |
are clearly discernible until |
13 DAP |
Arabidopsis thaliana |
| zou_nac seeds at 21 DAP |
were still plump with |
apparently viable endosperm |
Arabidopsis thaliana |
| Opaque-2 |
is involved in |
maize endosperm development |
Zea mays |
| triploid endosperm nuclei |
display |
15 centromeres marked by (CENH3, HTR12, AT1G01370) |
Arabidopsis thaliana |
| mRNA of the GluA-2 gene |
started to accumulate during the early stages of endosperm development |
|
|
| FIS-PRC2 mutants |
lack |
proper endosperm cellularization |
Arabidopsis thaliana |
| syncytial phase |
transitions to |
cellularized phase |
|
| (AGL62, AT5G60440) mutant |
shows |
precocious cellularization |
Arabidopsis thaliana |
| mn1 mutant |
showed |
reduced cell size during endosperm development |
Zea mays |
| metabolic physiology at 20 and 28 DAP |
is quite different between |
Mn1 and mn1 mutant |
Zea mays |
| INDUCER OF CBF EXPRESSION1 (ATICE1, ICE1, SCREAM, SCRM, AT3G26744) |
has strong expression in |
embryo-surrounding endosperm |
|
| TtPARC6 expression |
is lowest during |
late developmental stages |
Triticum turgidum |
| 11 SNPs in ta1-causal gene |
showed no changes in |
aleurone thickness |
Oryza sativa |
| interspecific cross between female Arabidopsis thaliana (4n) and male Arabidopsis arenosa (4n) |
shows |
delayed timing of cellularization |
Arabidopsis thaliana; Arabidopsis arenosa |
| endosperm cells in 4n self-pollinated plants |
are comparable in size to |
endosperm cells in 4n × 2n cross |
Oryza sativa |
| CpG DNA methylation at the 3′ regulatory region of the (AGL37, PHE1, AT1G65330) locus |
is decreased in |
(EMB173, FIS1, MEA, SDG5, AT1G02580) mutants |
Arabidopsis thaliana |
| (AtTRM61, TRM61, AT5G14600) |
may be required for |
endosperm development |
Arabidopsis thaliana |
| HAIKU2 (IKU2, AT3G19700) |
participates in |
endosperm development pathway |
Arabidopsis thaliana |
| large nuclei surrounded by dense cytoplasm in zou_nac |
suggests |
change in endosperm nature |
Arabidopsis thaliana |
| endosperm development |
follows |
nuclear-type program |
Arabidopsis thaliana |
| OsmtSSB1-mediated mitochondrial function |
plays a critical role in |
subaleurone cell-fate determination |
Oryza sativa |
| central vacuole in endosperm |
is replaced by |
endosperm cells by 4 DAP |
Oryza sativa |
| PcG-dependent histone methylation |
may be sufficient to repress |
autonomous central cell proliferation |
Arabidopsis thaliana |
| AtRFC4 loss of function |
caused |
endosperm with six to eight free nuclei |
Arabidopsis thaliana |
| rfc4-1/+ complementation with AtRFC4 expression driven through FIS2pro |
could not completely restore |
defective endosperm |
Arabidopsis thaliana |
| 2n self-pollinated seeds |
show |
endosperm nuclei increase at similar rates between 1.0 and 2.0 DAP |
Oryza sativa |
| 738 ± 31 endosperm nuclei in 2n × 4n cross at 1.5 DAP |
is approximately |
166% of that in 4n self-pollinated plants |
Oryza sativa |
| fertilized central cell |
undergoes |
endoreduplication |
|
| DEMETER (DME, EMB1649, AT5G04560) |
removes DNA methylation marks specifically in |
central cell |
Arabidopsis thaliana |
| pathways mediating non-CpG methylation in seeds |
are dependent on |
a decrease in CpG methylation |
Arabidopsis thaliana |
| interaction network and coexpression of AGL genes |
consistent with |
shared function during early endosperm development |
Arabidopsis thaliana |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) -1 heterozygous plants |
show lower frequency of |
autonomous endosperm development |
Arabidopsis thaliana |
| fis class mutants |
initiate central cell proliferation and endosperm development in the absence of fertilization |
central cell proliferation and endosperm development |
Arabidopsis thaliana |
| ZmMRP-1 |
is specifically expressed in |
transfer cell layer of the maize endosperm |
Zea mays |
| EMBRYO SURROUNDING FACTOR 1 (ESF1) peptides |
accumulate in |
embryo-surrounding endosperm cells |
Arabidopsis thaliana |
| changes in gene expression in endosperm |
may alter |
endosperm development |
|
| INDUCER OF CBF EXPRESSION1 (ATICE1, ICE1, SCREAM, SCRM, AT3G26744) |
is expressed in |
testa |
|
| genetic and physiological factors |
result in modulation of |
rhamnogalacturonan I (RG-I) structure |
|
| pGCD1::GCD1_EGFP construct |
shows GCD1 protein expression in |
early endosperm at two endosperm nucleus stage |
Arabidopsis thaliana |
| point mutation in junction between intron 4 and exon 5 of OsmtSSB1 |
caused |
(ASD, ATA1, TA1, AT3G42960) phenotypes |
Oryza sativa |
| subaleurone cells in rice endosperm |
differentiate to |
starchy endosperm at late stage of endosperm development |
Oryza sativa |
| endosperm from interploidy cross 4x O. sativa × 2x O. sativa |
cellularization was observed precociously at |
2 DAP |
Oryza sativa |
| cellularization timing in ploidy manipulation cross |
is similar to |
cellularization timing in self-pollinated diploid plants |
Oryza sativa; Oryza longistaminata |
| initial syncytial development followed by cellularization |
is |
conserved |
Oryza sativa; Arabidopsis thaliana |
| Opaque2 |
is |
known regulator of zein expression or endosperm development |
Zea mays |
| weak TIP3-GFP signal in starchy endosperm cells |
indicated |
gradual loss of TIP3-GFP-labelled membranes |
Hordeum vulgare |
| protein storage vacuoles (PSVs) in the central starchy endosperm |
do not appear to undergo |
fusion |
Hordeum vulgare |
| hybridization barrier |
may result in |
enlarged and swollen endosperm |
|
| paternal genome excess in Oryza sativa |
results in |
increased rate of nuclear division |
Oryza sativa |
| interspecific crosses |
may share common outcomes with |
interploidy crosses |
|
| 2n × 4n cross |
produces seeds filled with |
viscous liquid at 7 DAP |
Oryza sativa |
| 2n × 4n cross seeds |
show |
syncytium and central vacuole persisted to 7 DAP |
Oryza sativa |
| ovules containing autonomous endosperm |
express |
KS22 marker |
|
| pGCD1::GCD1_EGFP construct |
shows GCD1 protein expression in |
early endosperm at four endosperm nucleus stage |
Arabidopsis thaliana |
| efficient energy supply |
is critical for |
formation of single-layered aleurone in rice endosperm |
Oryza sativa |
| endosperm of (ASG6, CRK2, AT1G70520) |
was |
morphologically abnormal |
Zea mays |
| three TaNAC77 homoeologs |
shared |
similar expression profile |
Triticum aestivum |
| genomic cytosine methylation |
could impact |
accumulation of endosperm storage compounds |
Zea mays |
| repression of expression of maternal alleles of paternally-expressed genes (PEGs) |
occurs throughout |
endosperm development |
|
| reciprocal cross between female diploid Arabidopsis thaliana (2n) and male tetraploid Arabidopsis thaliana (4n) with paternal genome excess |
shows |
increased rate of mitotic division |
Arabidopsis thaliana |
| OsMADS87 gene |
is |
maternally expressed imprinted gene |
Oryza sativa |
| global DNA demethylation |
may affect many genes |
many genes |
Arabidopsis thaliana |
| (DDM2, DMT01, DMT1, MET1, MET2, METI, AT5G49160) |
potentially interacts with |
(FIE, FIE1, FIS3, AT3G20740) |
Arabidopsis thaliana |
| transcriptome analysis |
showed |
sharp transition at 9–10 days after pollination |
Zea mays |
| genetic networks |
regulate |
cell cycle regulation |
Oryza sativa |
| Helitrons flanking imprinted and non-imprinted genes |
are important regulators of |
endosperm development |
|
| rice endosperm |
continues to proliferate and to support |
seedling growth |
Oryza sativa |
| DNA methylation and H3K27me3 marks |
are mostly mutually exclusive |
H3K27me3 targets in the endosperm |
Arabidopsis thaliana |
| DNA methylation in non-CpG contexts |
is slightly increased at |
both (AGL37, PHE1, AT1G65330) and (EMB173, FIS1, MEA, SDG5, AT1G02580) loci |
Arabidopsis thaliana |
| a decrease in CpG methylation and increase in non-CpG methylation |
is observed in |
(EMB173, FIS1, MEA, SDG5, AT1G02580) mutant background for two target loci |
Arabidopsis thaliana |
| (DME, EMB1649, AT5G04560) |
is an antagonist of |
(DDM2, DMT01, DMT1, MET1, MET2, METI, AT5G49160) |
Arabidopsis thaliana |
| inhibition effect of the three SBEs in TRS endosperm development process |
was gradually enhanced |
|
|
| FIS-PRC2 complex |
likely functions through their 5′-flanking sequences to down-regulate |
actively expressed AGL genes prior to endosperm cellularization irrespective of their imprinting status |
Arabidopsis thaliana |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) |
has role in inhibition of |
endosperm proliferation |
Arabidopsis thaliana |
| over-proliferation phenotype caused by (EMB173, FIS1, MEA, SDG5, AT1G02580) mutation |
is suppressed in |
transgenic plants expressing antisense strand of PHERES1 (AGL37, PHE1, AT1G65330) gene driven by (EMB173, FIS1, MEA, SDG5, AT1G02580) promoter |
Arabidopsis thaliana |
| C2.2 genes |
maintained |
high relative levels of mRNA at 4 to 5 DAP |
Arabidopsis thaliana |
| (AGL28, AT1G01530) |
showed |
more delayed down-regulation pattern |
Arabidopsis thaliana |
| parent-specific activity of promoter and/or full-length gene fusions for 15 C2 genes |
assayed in |
fis2-8 and mea-3 mutant backgrounds at 2 and 5 DAP |
Arabidopsis thaliana |
| FIS-PRC2 |
required to repress expression of |
majority of C2 genes |
|
| C2 genes |
expressed in |
coenocytic endosperm |
|
| mid phase of endosperm development |
is |
6–15 DAP |
Zea mays |
| mitotic divisions |
give rise to |
most of the cells comprising the starchy endosperm |
Zea mays |
| cell number in wild type and OE lines |
showed no significant difference |
between wild type and OE lines |
Zea mays |
| Bdiku2 mutant 4-11 |
had begun to cellularize around |
few nuclei in embryo-surrounding region at 2 DAP |
Brachypodium distachyon |
| FIS-PRC2 |
mediates generalized repression of |
C2 subclade of type I MADS-box genes |
Arabidopsis thaliana |
| (AGL96, AT5G06500) |
showed |
more delayed down-regulation pattern |
Arabidopsis thaliana |
| antagonistic roles of parental chromosomes |
explains |
opposite phenotypes in endosperm |
|
| MEDEA (EMB173, FIS1, MEA, SDG5, AT1G02580) and FERTILIZATION INDEPENDENT SEED2 (FIE2, FIS2, AT2G35670) |
regulate endosperm development in opposite directions to |
PHERES1 (AGL37, PHE1, AT1G65330) |
Arabidopsis thaliana |
| balance of activity between imprinted genes |
is expected to be tightly regulated |
endosperm development |
|
| mis-expression of the target genes |
would cause |
endosperm abnormality |
Arabidopsis thaliana; Arabidopsis arenosa |
| endosperm from ploidy manipulation cross 4x O. sativa × 2x O. longistaminata |
was completely cellularized by |
3 DAP |
Oryza sativa; Oryza longistaminata |
| increased cell area during cellular stage |
might be the result of |
increased vacuolar volume and/or endoreduplication cycle |
Oryza sativa |
| Parentally biased expression |
might achieve |
optimized stoichiometry of duplicated ERFs that are crucial for the endosperm developmental program |
|
| (RGE1, ZOU, AT1G49770) mutants |
retain |
excess of endosperm material at maturity |
|
| nonmutagenized QPM |
accumulated smaller protein bodies |
protein bodies 30% to 40% of W64A wild-type size |
Zea mays |
| T-DNA insertion lines |
do not show |
autonomous endosperm development |
Oryza sativa |
| (FIE, FIE1, FIS3, AT3G20740) mutants |
identified |
PHERES1 (AGL37, PHE1, AT1G65330) |
Arabidopsis thaliana |
| FERTILIZATION-INDEPENDENT SEED (FIS)-Polycomb Repressive Complex2 (PRC2) |
is |
putative H3K27 methyltransferase |
Arabidopsis thaliana |
| (AGL35, AT5G26630) |
showed |
more delayed down-regulation pattern |
Arabidopsis thaliana |
| cellularization timing in endosperm |
is opposite in |
reciprocal cross |
|
| (AGL62, AT5G60440) and (AGL37, PHE1, AT1G65330) MADS-box heterodimer complex |
may regulate |
endosperm cellularization |
Arabidopsis thaliana |
| FIS-PRC2 complex |
maintains |
maternal but not the paternal imprinting of the imprinted AGL genes |
Arabidopsis thaliana |
| H3K27me3 modifications at FIS-PRC2-regulated C2 genes |
undergo dynamic change in pattern during |
early endosperm development |
|
| cellularization |
extends across |
peripheral domain toward chalazal domain |
Arabidopsis thaliana |
| (AGL62, AT5G60440) mutant |
exhibits |
precocious cellularization |
Arabidopsis thaliana |
| imprinting of rice and maize (FIE, FIE1, FIS3, AT3G20740) genes |
may indicate |
similar function in endosperm development |
Oryza sativa; Zea mays |
| ZmFie1 and ZmFie2 RNAi lines |
have not shown |
autonomous endosperm development |
Zea mays |
| FERTILIZATION INDEPENDENT ENDOSPERM (FIE, FIE1, FIS3, AT3G20740) |
is |
component of PRC2 Polycomb protein complex |
Arabidopsis thaliana |
| redundant functions in (AGL37, PHE1, AT1G65330) family |
may explain |
lack of obvious phenotype in (AGL37, PHE1, AT1G65330) antisense transgenic plants without (EMB173, FIS1, MEA, SDG5, AT1G02580) mutation |
Arabidopsis thaliana |
| 10 lines with fully viable opaque kernels |
were candidates for factors specifically involved in |
o2 endosperm modification and/or vitreous endosperm formation |
Zea mays |
| cellularization |
begins after |
eighth mitosis |
Arabidopsis thaliana |
| PHERES1 (AGL37, PHE1, AT1G65330) |
is |
type-I MADS-box transcription factor |
Arabidopsis thaliana |
| global DNA hypomethylation |
causes |
prolonged coenocytic phase |
Arabidopsis thaliana |
| (AGL62, AT5G60440) binding to C2 proteins |
supports notion that |
at least some C2 genes regulate endosperm development |
|
| delayed cellularization |
is correlated with |
larger seed size |
|
| endosperm abnormality |
may be predicted using |
endosperm balance number |
|
| interspecific cross between female Arabidopsis thaliana (4n) and male Arabidopsis arenosa (4n) |
shows |
decreased rate of mitotic division |
Arabidopsis thaliana; Arabidopsis arenosa |
| synergistic effect of DNA methylation and H3K27me3 |
is probably responsible for the enhancement of |
autonomous endosperm development in (DDM2, DMT01, DMT1, MET1, MET2, METI, AT5G49160) -3 ; (EMB173, FIS1, MEA, SDG5, AT1G02580) -1 plants |
Arabidopsis thaliana |
| (DDM2, DMT01, DMT1, MET1, MET2, METI, AT5G49160) |
does not have a major effect on repression of |
(EMB173, FIS1, MEA, SDG5, AT1G02580) before fertilization |
Arabidopsis thaliana |
| ATHILA retrotransposons |
misexpression of |
endosperm development |
Arabidopsis thaliana; Arabidopsis arenosa |
| endosperm of O. sativa × O. longistaminata cross |
showed abnormal |
cell size |
Oryza sativa; Oryza longistaminata |
| imprinting status of MADS87 |
correlated with |
abnormal or restored endosperm phenotype |
Oryza sativa; Oryza longistaminata |
| FIS-PRC2 |
regulates |
C2 subclade of type I MADS-box genes |
Arabidopsis thaliana |
| autonomous development of endosperm |
may be controlled by |
other mechanisms yet to be identified |
Oryza sativa |
| PHERES1 (AGL37, PHE1, AT1G65330) |
is |
up-regulated in (EMB173, FIS1, MEA, SDG5, AT1G02580) and (FIE, FIE1, FIS3, AT3G20740) mutants |
Arabidopsis thaliana |
| C2 genes (with exception of MEGs (AGL36, AT5G26650) (AGL90, AT5G27960) and (AGL96, AT5G06500) and PEGs (AGL23, AT1G65360) and (AGL37, PHE1, AT1G65330) ) |
generally exhibited |
biallelic expression pattern during coenocytic endosperm development |
Arabidopsis thaliana |
| FIS-PRC2 complex |
has dual role in |
regulation of AGL genes |
Arabidopsis thaliana |
| FIS-PRC2 complex |
may remain associated with |
C2 loci in form of bivalent chromatin domains |
|
| Ploidy barrier to endosperm model |
could apply similar mechanism to |
imprinted PcG gene regulation of endosperm development |
|
| AGAMOUS-LIKE62 (AGL62, AT5G60440) |
regulates |
seed size |
Arabidopsis thaliana |
| (EMB173, FIS1, MEA, SDG5, AT1G02580) mutants |
identified |
PHERES1 (AGL37, PHE1, AT1G65330) |
Arabidopsis thaliana |
| correlation between cell number and endosperm thickness |
was strong in early endosperm development but became less evident during endosperm filling |
endosperm development stages |
Zea mays |
| increased endosperm cell ploidy |
is associated with |
cell size |
Zea mays |
| defective seeds from pBdIKU2 :: BdIKU2 transgenic plants |
had |
uncellularized or partially cellularized endosperm |
Arabidopsis thaliana |
| cellularization in Bdiku2 mutant 4-11 |
progressed rapidly toward |
central vacuoles from 3 to 4 DAP |
Brachypodium distachyon |
| FIS-PRC2 complex |
may reassociate with |
C2 loci during endosperm cellularization |
|
| (EMB173, FIS1, MEA, SDG5, AT1G02580) female gametophytes |
develop into |
seed-like structures with diploid endosperms |
Arabidopsis thaliana |
| changes in patterns of expression of imprinted genes |
may disrupt balance and cause |
endosperm abortion |
|
| interspecific hybrid between Arabidopsis thaliana and Arabidopsis arenosa |
shows |
delayed cellularization in the endosperm |
Arabidopsis thaliana; Arabidopsis arenosa |
| majority of the C2 genes tested (including (AGL48, AT2G40210) and (AGL91, AT3G66656) ) |
were |
biallelically expressed as promoter reporters |
Arabidopsis thaliana |
| hybrid endosperm |
shows |
overgrowth with altered or arrested embryo development |
Arabidopsis thaliana; Arabidopsis arenosa |
| 50-kD γ-zein |
does not appear to be involved in |
protein body initiation |
Zea mays |
| type I MADS-box transcription factor genes |
may constitute |
regulatory circuitry linking epigenetic mechanisms with regulation of genes expressed during early endosperm development |
Arabidopsis thaliana |
| FIS-PRC2 regulation of early endosperm development |
has |
dual role |
Arabidopsis thaliana |
| generalized repression of C2 subclade by FIS-PRC2 |
is associated with |
endosperm cellularization |
Arabidopsis thaliana |
| abnormal development |
occurs in |
hybrid endosperm |
|
| paternal alleles of MEGs (AGL36, AT5G26650) (AGL90, AT5G27960) and (AGL96, AT5G06500) |
remained |
silenced at 2 and 5 DAP |
Arabidopsis thaliana |
| seed size |
is positively associated with |
extent of coenocytic endosperm development |
Arabidopsis thaliana |
| observed up-regulation of the C2 genes at 5 DAP |
may result from |
extended coenocytic phase |
Arabidopsis thaliana |
| Polar Nuclei Activation model |
could apply similar mechanism to |
imprinted PcG gene regulation of endosperm development |
|
| line 107 protein bodies |
were identified as being similar in size to |
wild type protein bodies |
Zea mays |
| coenocytic phase |
is characterized by |
series of mitoses without cytokinesis |
Arabidopsis thaliana |
| coenocytic endosperm |
differentiates into |
peripheral domain |
Arabidopsis thaliana |
| 5′-flanking sequences of (AGL37, PHE1, AT1G65330) |
are sufficient to mediate |
FIS-PRC2-dependent repression of either allele during the transition to cellularization |
Arabidopsis thaliana |
| Polar Nuclei Activation hypothesis |
was proposed to explain |
endosperm incompatibility in oats |
Avena sativa |
| antagonistic regulation of endosperm development by (EMB173, FIS1, MEA, SDG5, AT1G02580) (FIE2, FIS2, AT2G35670) and (AGL37, PHE1, AT1G65330) |
explains |
observed phenotypes of interspecific crosses |
Arabidopsis thaliana |
| O2-ZmGRAS11-centered transcriptional regulatory network |
coordinately regulates |
endosperm filling and cell expansion |
Zea mays |
| coupled regulation of cell expansion and endosperm filling |
is advantageous in that |
storage capacity and biosynthetic capability are coordinated to maximize cell volume and contents on real-time basis |
Zea mays |
| cellularization in Atiku2 |
was completed by |
5 DAP |
Arabidopsis thaliana |
| number of syncytial nuclei at 1.0, 1.5 and 2.0 DAP in self-pollinated 2x O. sativa |
showed no significant difference compared to |
number of syncytial nuclei in interspecific cross 2x O. sativa × 2x O. longistaminata |
Oryza sativa; Oryza longistaminata |
| endosperm in A. thaliana |
is |
ephemeral and degenerates into single layer at maturity |
Arabidopsis thaliana |
| reduced expression of OsMADS87 |
is correlated with |
precocious endosperm cellularization |
Oryza sativa |
| almost half of the DEGs |
were upregulated in |
each transition between developmental stages |
Triticum urartu |
| DNA methylation |
plays crucial role in |
key biological events during maize endosperm development |
Zea mays |
| endosperm development of T. urartu |
shows decrease in |
fraction of medium- and high-prevalence genes |
Triticum urartu |
| RNA-seq |
revealed |
crucial roles of genetic controls in rice endosperm development |
Oryza sativa |
| RNA-directed DNA methylation (RdDM) |
might be important for mediating balance between |
maternal and paternal contributions to the endosperm |
Arabidopsis thaliana |
| nonmutagenized QPM |
accumulated protein bodies in higher number than |
wild type |
Zea mays |
| type I MADS-box genes |
are regulated by |
polymerase IV-dependent small interfering RNAs (p4-siRNAs) |
Arabidopsis thaliana |
| C1 genes |
showed higher relative expression levels either before fertilization or upon |
endosperm cellularization (4 and 5 DAP) |
Arabidopsis thaliana |
| FIS-PRC2 |
is likely required for |
proper down-regulation of C2 genes at the onset of endosperm cellularization |
Arabidopsis thaliana |
| all three homeologs |
show the same pattern through |
endosperm development |
Triticum aestivum |
| imprinting status of YUCCA11 |
correlated with |
abnormal or restored endosperm phenotype |
Oryza sativa; Oryza longistaminata |
| endosperm abnormality |
is probable cause of |
effective hybridization barrier |
Oryza sativa |
| disruption of any AtMCM2–7 gene |
results in |
enlarged endosperm free nuclei |
Arabidopsis thaliana |
| deletion mutagenesis in maize |
has potential as tool for investigating |
endosperm maturation |
Zea mays |
| histone modification |
likely contributes to |
epigenetic mechanisms regulating endosperm development |
Arabidopsis thaliana |
| (AGL91, AT3G66656) and (AGL33, AT2G26320) |
were not observed to be up-regulated in |
mutants at 5 DAP |
Arabidopsis thaliana |
| (AGL62, AT5G60440) |
is expressed in |
endosperm during syncytial phase |
Arabidopsis thaliana |
| aborted ovules in rfc4-1/+ and rfc4-2/+ |
have far fewer |
endosperm free nuclei than wild type |
Arabidopsis thaliana |
| endosperm undergrowth |
is |
type of developmental abnormality in endosperm |
|
| emp8 mutants |
show severely arrested |
endosperm development |
Zea mays |