| knockout of (ACBP3, AT4G24230) |
caused adverse impact on |
anther development in Ler-0 |
Arabidopsis thaliana |
| OsALKBH5 protein levels in anthers |
tended to stay high after |
stage 10 |
Oryza sativa L. ssp. japonica |
| development of the hvtdf1-2 dPMCs |
progressed at a similar pace as in |
WT PMC stage |
Hordeum vulgare |
| polymorphism in the regulatory region of (ACBP3, AT4G24230) |
enabled |
(ACBP3, AT4G24230) function in anther development in Ler-0 |
Arabidopsis thaliana |
| upregulation of (NZZ, SPL, AT4G27330) |
failed to rescue |
acbp3-2 phenotype |
Arabidopsis thaliana |
| (AtC3H15, CDM1, AT1G68200) tapetum morphology |
appeared normal and different from |
known mutants with defective tapetum (e.g. (DYT1, AT4G21330) and (AMS, AT2G16910) ) |
Arabidopsis thaliana |
| Tapetal cell number increase in hvtdf1 mutant |
occurred at |
Meiosis I to Microspore release stage |
Hordeum vulgare |
| differential developmental progress observed between the abaxial and the adaxial lobes in (bZIP21, TGA9, AT1G08320) (bZIP65, TGA10, AT5G06839) anthers |
indicates that |
four lobes do not develop simultaneously |
Arabidopsis thaliana |
| highly-expressed transcripts and proteins of (ACBP3, AT4G24230) in Ler-0 anthers |
promote |
anther development |
Arabidopsis thaliana |
| delayed expression in most pathways in hvtdf1 |
corresponds to |
phenotype observed from anther sections |
Hordeum vulgare |
| differences in the regulatory network governing anther development between Ler-0 and Col-0 |
exist between |
Ler-0 and Col-0 |
Arabidopsis thaliana |
| hvtdf1-2 plants |
had |
smaller anthers compared to WT |
Hordeum vulgare |
| meiosis process of PMCs |
is not independent |
it requires communication between germline cells and the surrounding tapetal cells |
Hordeum vulgare |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) mutants in Col-0 |
is similar to |
acbp3-2 mutant in Ler-0 |
Arabidopsis thaliana |
| genetic polymorphism in the regulatory region of (ACBP3, AT4G24230) |
enables |
downstream function of (ACBP3, AT4G24230) to be regulated by (NZZ, SPL, AT4G27330) |
Arabidopsis thaliana |
| meiosis |
could not be observed in |
hvtdf1-2 dPMCs |
Hordeum vulgare |
| OsALKBH5 mRNA in anthers |
generally increased until Stage 10 and then decreased |
OsALKBH5 mRNA abundance |
Oryza sativa L. ssp. japonica |
| WT tapetum cells and meiotic microspore cells (MMCs) |
were deeply stained using 0.05% toluidine blue solution |
indicating active biological processes |
Hordeum vulgare |
| anther primordium |
forms from |
floral meristem |
|
| OsERS1 transcript levels in anther |
are relatively high during |
stages 3 to 6 |
Oryza sativa |
| OsFBK1 overexpression transgenics |
showed |
aberrant appearances of ridged anther cuticle |
Oryza sativa |
| Attdf1 mutant |
showed |
extensive increased tapetal cell divisions |
Arabidopsis thaliana |
| Genes from Cluster B |
started expression at stage 7 and peaked at 8a2, then decreased at stage 8b |
indicating stage-specific expression |
Hordeum vulgare |
| rescued anther phenotype in acbp3-2 × La-0 |
supports |
hypothesis of (ACBP3, AT4G24230) function in Ler-0 and ERECTA signaling pathway relationship |
Arabidopsis thaliana |
| Spikes from the hvtdf1-2 mutant |
were collected from |
stage 8a2 to 8b with two biological replicates |
Hordeum vulgare |
| acbp3-1 × er-1 |
did not exhibit the same phenotype as |
acbp3-2 |
Arabidopsis thaliana |
| polymorphism within the (ACBP3, AT4G24230) promoter |
enabled |
(NZZ, SPL, AT4G27330) function in anther development |
Arabidopsis thaliana |
| no obvious tetrads |
could be observed in |
mutant line |
Hordeum vulgare |
| tapetum layer |
shows least ROS accumulation |
anther development stage 5 |
|
| observed defect for acbp3-2 in sporocyte formation within adaxial lobes |
supports |
regulatory role of (NZZ, SPL, AT4G27330) on (ACBP3, AT4G24230) expression in Ler-0 anthers |
Arabidopsis thaliana |
| knockout mutants of six BZR genes |
show |
abnormal sporocyte formation |
Arabidopsis thaliana |
| differential impact of (ACBP3, AT4G24230) on abaxial and adaxial lobe development observed in -2 anthers |
imply that |
(ACBP3, AT4G24230) likely functions downstream of (NZZ, SPL, AT4G27330) in Ler-0 |
Arabidopsis thaliana |
| WT tapetum cells at tetrad stage |
stained much darker and started |
the condensation process with tetrads attached to the inner tapetum surface |
Hordeum vulgare |
| primary sporogenous cell layer in staged-3 anthers of (NZZ, SPL, AT4G27330) |
failed to form |
microsporocytes |
Arabidopsis thaliana |
| these results |
may explain |
the observed delayed endomitosis and irregular meiosis phenotype |
Hordeum vulgare |
| (NZZ, SPL, AT4G27330) |
has well-established role in |
sporocyte formation |
Arabidopsis thaliana |
| AtVRLK1 overexpression (OE) plants |
show severe sterility with |
anther dehiscence failure |
Arabidopsis thaliana |
| GRXs |
regulate |
differentiation of microsporocytes |
Oryza sativa; Zea mays; Arabidopsis thaliana |
| OsFBK1 |
plays role in |
development of rice anthers |
Oryza sativa |
| reported exclusion of (MIR319, MIR319B, AT5G41663) from late anther development |
explains |
absence of (MIR319, MIR319B, AT5G41663) from pollen |
|
| comparison between hvtdf1-2 stage 8b vs WT stage 8b |
was performed |
to control for developmental stage differences |
Hordeum vulgare |
| recovery of normal anthers in acbp3-2 × La-0 plants |
substantiates that |
(ACBP3, AT4G24230) function is related to the ER-mediated signaling pathway |
Arabidopsis thaliana |
| (NZZ, SPL, AT4G27330) function |
differs in |
Col-0 and Ler-0 ecotypes |
Arabidopsis thaliana |
| anther development stage 5 |
all four somatic cell layers are established |
anther development |
|
| ACBP3pro (Ler)::ACBP3::GFP |
rescued |
aberrant adaxial anther lobe formation |
Arabidopsis thaliana |
| anthers in COM-L flowers |
appear |
normal |
Arabidopsis thaliana |
| distinct function of (ACBP3, AT4G24230) in Ler-0 |
is not solely attributed to |
er mutation |
Arabidopsis thaliana |
| delay in development |
is supported by |
both the morphology and transcriptome data |
Hordeum vulgare |
| ERECTA |
modulates |
(ACBP3, AT4G24230) function differently in Landsberg and Columbia |
Arabidopsis thaliana |
| delayed pathways |
endorses |
retarded anther morphology |
Hordeum vulgare L. |
| WT during microspore development stage |
showed |
vacuoles in the microspores increased in size, resulting in round, mature microspores with off-set nuclei |
Hordeum vulgare |
| comparison of the transcriptome changes, particularly downregulated genes (q < 0.05 and Fold change < −1) |
was performed between |
the barley and Arabidopsis (ATMYB35, MYB35, TDF1, AT3G28470) mutants |
Hordeum vulgare; Arabidopsis thaliana |
| proper (ACBP3, AT4G24230) expression |
is crucial for |
anther development in Ler-0 |
Arabidopsis thaliana |
| (ATMYB35, MYB35, TDF1, AT3G28470) and (AMS, AT2G16910) |
each takes responsibility to regulate |
key anther developmental processes |
Hordeum vulgare; Arabidopsis thaliana |
| COM-C plants harboring ACBP3pro (Col)::ACBP3::GFP |
were used to complement |
acbp3-2 mutant |
Arabidopsis thaliana |
| abnormal (ACBP3, AT4G24230) expression |
adversely affects |
anther development |
Arabidopsis thaliana |
| hvtdf1-2 tapetum cells |
did not show detectable |
endomitosis |
Hordeum vulgare |
| WT tapetum layer during microspore development stage |
started degeneration and became |
thinner |
Hordeum vulgare |
| homozygous Arabidopsis (ATMYB35, MYB35, TDF1, AT3G28470) mutant plants carrying the HvTDF1 gene |
showed recovered fertility as evidenced by |
elongated siliques, fully formed dehiscent anthers and viable pollen |
Arabidopsis thaliana |
| barley homozygous mutant plants |
had |
no seed filling in mutant spikes |
Hordeum vulgare |
| tapetal cell numbers on independent transverse sections |
were counted in |
hvtdf1-2 mutant and the WT |
Hordeum vulgare |
| ACBP3pro (Ler)::ACBP3::GFP |
showed high expression in |
anthers |
Arabidopsis thaliana |
| double mutant of (bZIP21, TGA9, AT1G08320) and (bZIP65, TGA10, AT5G06839) |
exhibit |
normal abaxial anther lobes but variable or disorganized adaxial lobes |
Arabidopsis thaliana |
| differential anther morphology observed between acbp3-1 × er-1 and acbp3-2 plants |
is consistent with |
differential acyl-CoA content between acbp3-1 × er-1 and acbp3-2 plants |
Arabidopsis thaliana |
| HvTDF1 |
starts to express in spikes from |
Pollen Mother Cell (PMC) stage until the tetrad stage |
Hordeum vulgare |
| development process in the hvtdf1 mutant |
appears delayed by almost one stage compared with |
WT |
Hordeum vulgare |
| ROS accumulation in tapetum layer |
starts from |
anther meiosis initiation |
|
| ACBP3pro (Ler)::ACBP3 |
restored |
anther development in acbp3-2 |
Arabidopsis thaliana |
| ACBP3pro (Col)::ACBP3 |
did not restore |
(ACBP3, AT4G24230) expression in -2 |
Arabidopsis thaliana |
| (NZZ, SPL, AT4G27330) interaction with the AT~TATA Box |
influences |
anther development and acyl-CoA homeostasis |
Arabidopsis thaliana |
| SPOROCYTELESS/NOZZLE ( (NZZ, SPL, AT4G27330) ) |
play pivotal roles in |
specification of archesporial cell fate and delineation of archesporial and somatic cells |
Arabidopsis thaliana |
| GRF-GIF duo |
is |
novel positive regulator in specifying archesporial cells |
Arabidopsis thaliana |
| loss of function of OsERS1 |
causes |
abnormal anther morphogenesis |
Oryza sativa |
| OsCCR14 |
could play important roles in |
lignification in anthers |
Oryza sativa |
| OsROXY2 |
is detectable in |
tetrads |
Oryza sativa |
| ERECTA-related role of (ACBP3, AT4G24230) |
is not required in |
Col-0 |
Arabidopsis thaliana |
| osers1 anther at stage 3 |
has PPCs organized in |
irregular mosaic pattern |
Oryza sativa |
| comparison of transcriptome data between the hvtdf1 mutant and WT |
was conducted to understand |
the biological function of HvTDF1 in barley anther development |
Hordeum vulgare |
| consistent high expression level of (AMS, AT2G16910) from the anther development stage 7–8b |
is critical for |
normal anther development |
Hordeum vulgare |
| wild-type barley |
shows equivalent progression of anther development until |
stage 7 |
Hordeum vulgare |
| somatic and germline cell layers in anther |
show specific dynamic hypoxia and ROS production in |
each cell layer |
|
| enrichment of highly active metabolic processes in (ATMYB35, MYB35, TDF1, AT3G28470) |
suggests anther status is still under |
developmental flux when (ATMYB35, MYB35, TDF1, AT3G28470) is expressed |
Arabidopsis thaliana |
| normal anthers in acbp3-1 × er-1 |
support that |
(ACBP3, AT4G24230) did not function through the ER-mediated signaling pathway in Col-0 |
Arabidopsis thaliana |
| osers1 anther wall layers in later stages |
had stronger H2O2 signals compared with |
wild-type anther wall layers |
Oryza sativa |
| high-temperature injury to anther development |
is correlated with |
depletion of the phytohormone auxin |
Hordeum vulgare; Arabidopsis thaliana |
| cell surface-localized leucine-rich repeat receptor-like kinases (LRR-RLKs) |
function in |
determining early anther cell fate |
Oryza sativa; Arabidopsis thaliana |
| cool-weather damage in rice |
causes |
abnormal abortion of developing pollen |
Oryza sativa |
| L2 layer |
undergoes rapid cell division at stage 2 and generates |
L2-derived (L2-d) cells |
|
| transcription factors |
have critical functions in |
specifying cell lineage and the fate of germ and somatic cells |
Arabidopsis thaliana; Oryza sativa; Zea mays |
| MIL1 expression |
shows no significant difference between |
wild-type and osers1 anthers |
Oryza sativa |
| TGA transcription factors |
function in |
anther development |
Arabidopsis thaliana |
| Ligands of LRR-RLKs |
function in |
determining early anther cell fate |
Oryza sativa; Arabidopsis thaliana |
| osers1 anthers |
have |
adaxial fused lobes |
Oryza sativa |
| changes in redox status and cell behavior |
leads to |
changes in anther development |
Oryza sativa |
| L1 layer forming a rigid enclosure |
aggravates |
disarranged cell layers |
Oryza sativa |
| ccc anthers |
failed to |
dehisce |
Arabidopsis thaliana |
| TaNAC69 genes |
is developmentally regulated in |
anthers |
Triticum aestivum |
| early and excessive autophagy and premature PCD of tapetal cells |
ultimately leading to |
metabolic reprogramming and interruption of anther development |
Zea mays |
| detailed mechanisms of sugar transport among four layers |
remain elusive |
anther development |
|
| SlIDA promoter-β-glucuronidase (GUS) fusion construct |
shows weak GUS activity in stamens at |
earliest stages of anther development |
Solanum lycopersicum |
| OsROXY2 |
is expressed in |
anthers |
Oryza sativa |
| hormones |
have critical functions in |
specifying cell lineage and the fate of germ and somatic cells |
Arabidopsis thaliana; Oryza sativa; Zea mays |
| GRXs |
trigger archesporial fate determination by controlling |
redox status |
Oryza sativa; Zea mays; Arabidopsis thaliana |
| archesporial cells |
might silence |
mitochondria |
Zea mays |
| enclosed niche in anther primordium |
provides |
hypoxic conditions for archesporial cells |
Zea mays |
| OsFBK1 overexpression line anthers |
did not show |
apparent size reduction |
Oryza sativa |
| hypoxia |
maintains |
proper division and cell fate determination |
Zea mays |
| untreated osers1 anther |
had |
139.3 L2-d cells and 18.5 sporogenous-like cells |
Oryza sativa |
| OsROXY1 |
is detectable in |
tetrads |
Oryza sativa |
| epidermis |
protects against |
external environmental stresses |
|
| SlIDA |
is expressed in |
septum |
Solanum lycopersicum |
| OsMTR1 |
is essential for |
degeneration of tapetal cells |
Oryza sativa |
| exogenous gibberellin (GA) application |
promotes |
normal tapetal cell degeneration |
Oryza sativa |
| rice MULTIPLE SPOROCYTE1 (MSP1) and TAPETAL DETERMINANT1-LIKE A (OsTDL1A)/MIL2 |
are suggested to exert their effects in early anther cell differentiation via modulating |
redox status |
Oryza sativa |
| MIL2 expression |
shows no significant difference between |
wild-type and osers1 anthers |
Oryza sativa |
| auxin |
negatively regulates |
endothecium lignification |
Arabidopsis thaliana |
| OsROXY2 |
is mainly restricted to |
tapetum |
Oryza sativa |
| (NZZ, SPL, AT4G27330) and BAMs |
act as positive and negative regulators, respectively |
specification of archesporial cell fate |
Arabidopsis thaliana |
| (NZZ, SPL, AT4G27330) mutant anther |
consists of |
only somatic connective cells, producing no archesporial cells |
Arabidopsis thaliana |
| wild-type anthers treated with water |
showed slight increase in |
number of sporogenous-like cells from 7.5 to 9.2 |
Oryza sativa |
| OsROXY1 |
is expressed in |
anthers |
Oryza sativa |
| (ROXY1, AT3G02000) |
is expressed in |
anther development |
Arabidopsis thaliana |
| (AN3, ATGIF1, GIF, GIF1, AT5G28640) /2 35S:MIR396 |
produces |
no archesporial cells |
Arabidopsis thaliana |
| OsERS1 transcript levels in anther |
decreased in |
later stages |
Oryza sativa |
| mutations of the GRX MIL1 |
lead to |
misidentified anther wall cell layers |
Oryza sativa |
| OsFBK1 knockdown line |
has increased |
anther length |
Oryza sativa |
| auxin perception mutants |
show precocious |
anther development |
|
| accumulation and stabilization of SLR1/DELLA proteins |
would result in |
arrest of sporogenous cell proliferation at early developmental stages |
Oryza sativa |
| each lobe of the anther |
internally develops |
microsporangium |
Arabidopsis thaliana |
| AtVRLK1 overexpression (OE) anthers |
fail to open |
anther dehiscence |
Arabidopsis thaliana |
| growth-generated hypoxic conditions |
induce |
archesporial cells |
Zea mays |
| osers1 anther at stage 3 |
produces |
∼10 archesporial-like cells |
Oryza sativa |
| BARELY ANY MERISTEM ( (BAM1, AT5G65700) and (BAM2, BMY9, AT4G00490) ) |
play pivotal roles in |
specification of archesporial cell fate and delineation of archesporial and somatic cells |
Arabidopsis thaliana |
| IAA concentration |
leads to peak and finally declining when |
endothecium thickening commences |
|
| some of the archesporial cells of the (AN3, ATGIF1, GIF, GIF1, AT5G28640) triple mutant |
survived to form |
pollen grains |
Arabidopsis thaliana |
| genomic fragment containing wild-type OsERS1 gene |
rescues |
male-sterile phenotype in osers1 mutant |
Oryza sativa |
| OsERS1 mRNA |
is detected in |
SPC and sporogenous cells of stage 4 |
Oryza sativa |
| (AtC3H15, CDM1, AT1G68200) |
shows reduced expression in |
(EMS1, EXS, AT5G07280) mutant anthers |
Arabidopsis thaliana |
| early phase of anther development |
is more susceptible to |
low temperature (LT) stress |
Oryza sativa; Triticum aestivum; Hordeum vulgare |
| changed redox status |
disturbs |
early cell division in rice anthers |
Oryza sativa |
| OsERS1 |
maintains |
cell patterning |
Oryza sativa |
| (AtMYB103, ATMYB80, MS188, MYB103, MYB80, AT5G56110) gene |
is important for |
late tapetum development |
Arabidopsis thaliana |
| disarranged cell layers and rigid L1 enclosure |
created |
abaxial fused lobes |
Oryza sativa |
| ROS molecule H2O2 |
is initially accumulated in |
middle layer at stage 5 |
Oryza sativa |
| disruption of redox status in wild-type anthers by injecting low concentrations of H2O2 |
resulted in |
mimic of the phenotype of osers1 mutant |
Oryza sativa |
| ccc triple mutant |
displays |
non-dehiscent anthers |
Arabidopsis thaliana |
| phosphorylation of SPOROCYTELESS (NZZ, SPL, AT4G27330) |
enhances |
(NZZ, SPL, AT4G27330) protein stability |
Arabidopsis thaliana |
| CR-slida knockout lines |
exhibit |
failure of anther dehiscence |
Solanum lycopersicum |
| early stages of anther development (stages 1–5) |
determine |
formation of anther cell layers |
|
| loss of function of OsERS1 |
results in |
disarranged anther wall cell layers |
Oryza sativa |
| anther dehiscence |
has been largely studied in |
Arabidopsis thaliana |
Arabidopsis thaliana |
| OsJAR1 |
has a role in controlling |
anther development and dehiscence |
Oryza sativa |
| same size spikes |
were collected from |
WT and hvtdf1-2 representing equivalent development stages |
Hordeum vulgare |
| (ACBP3, AT4G24230) |
is |
subject of promoter polymorphism between Col-0 and Ler-0 |
Arabidopsis thaliana |
| phenotypic differences in anther development |
involved |
regulatory polymorphism in the promoter region of (ACBP3, AT4G24230) between Ler-0 and Col-0 |
Arabidopsis thaliana |
| (ACBP3, AT4G24230) |
has not been reported to influence |
anther development in Col-0 |
Arabidopsis thaliana |
| SlIDA |
is expressed in |
tapetum |
Solanum lycopersicum |
| microsporangium |
consists of |
three outer concentric parietal layers |
Arabidopsis thaliana |
| early anther development |
is characterized by |
rapid cell division and fate specification |
|
| primary parietal cells (PPCs) |
form via asymmetric cell division at stage 4 |
endothecium and secondary parietal cells (SPCs) |
|
| OsERS1 |
plays |
independent role with known regulators in early anther development |
Oryza sativa |
| premature endothecium lignification |
results in |
precocious anther dehiscence |
|
| cleared anthers viewed under UV in confocal microscope |
showed differences in |
dispersal of U-shaped wall thickenings in endothecium |
Oryza sativa |
| (ROXY1, AT3G02000) |
has transiently overlapping expression domains with |
(ROXY2, AT5G14070) |
Arabidopsis thaliana |
| anti-159 flowers |
have anthers reduced in size with shorter filaments and smaller pollen sacs than |
wild-type flowers |
Arabidopsis thaliana |
| Cytochrome P450 (CYP450) |
is involved in |
biosynthesis of hydroxylated fatty acids |
|
| ZmMs33/ZmGPAT6 |
was highly expressed in |
tapetum at early anther developmental stages |
Zea mays |
| (NZZ, SPL, AT4G27330) protein stability |
is required for |
Arabidopsis anther development |
Arabidopsis thaliana |
| (ATGPAT6, GPAT6, AT2G38110) mutants |
display |
altered endoplasmic reticulum (ER) profiles in the tapetum |
Arabidopsis thaliana |
| OsCP1 downregulation |
caused |
delayed tapetum PCD |
Oryza sativa |
| reactive oxygen species (ROS) production |
is required for |
normal anther development |
|
| tapetum |
contributes to |
pollen coat deposition |
|
| OsMED14_1 |
is expressed in |
tapetum and developing microspores |
Oryza sativa |
| septum PCD |
is required for |
anther dehiscence |
|
| early and excessive autophagy, premature PCD, and metabolic reprogramming in tapetal cells |
finally arrests |
elongation and development of mutant anthers |
Zea mays |
| failure to induce a ROS burst before anther dehiscence |
resulted in |
defect in secondary wall lignification in the anthers |
Arabidopsis thaliana |
| chloroplasts in En cells of plant anthers |
have been reported in |
barley |
Hordeum vulgare |
| transient peaks in ROS production |
occur in |
developing rice, tomato and tobacco anthers |
Oryza sativa; Solanum lycopersicum; Nicotiana tabacum |
| tapetum |
contributes to |
microspore release |
|
| APETALA3 (AP3, ATAP3, AT3G54340) PISTILLATA (PI), and AGAMOUS (AG) |
regulate |
subset of genes that promote anther development |
Arabidopsis thaliana |
| endothecium cells |
are vacuolated during |
anther stages 7 and 8 |
|
| tapetal cells in er-105 erl1-2 erl2-1 triple mutant |
possessed |
large vacuoles |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– mutant |
play crucial roles in |
anther lobe formation |
|
| proper coordination between cells |
is important for |
robust anther development |
Arabidopsis thaliana |
| Arabidopsis Tapetal Development and Function1 (ATMYB35, MYB35, TDF1, AT3G28470) |
is highly expressed in |
microspores |
Arabidopsis thaliana |
| (FAR2, MS2, AT3G11980) mutant |
causes |
shrivelled anthers |
Arabidopsis thaliana |
| apical and basal pore lengths |
shows significant differences among |
three rice genotypes |
Oryza sativa L. |
| ZmMs33 / ZmGPAT6 |
is critical for |
cell proliferation and expansion at late stages |
Zea mays |
| ZmMs33 / ZmGPAT6 |
is required for |
normal structure and function of En chloroplasts |
Zea mays |
| middle layer |
transports calcium ions to |
tapetum |
|
| dynamic changes in thickness of En and tapetum in WT and mutant anthers during stages 7–11 |
described |
possible coordinated relationship between En and tapetum |
Zea mays |
| GUS activity |
decreases with maturity and vanishes in |
mature anther and pollen grains |
Oryza sativa |
| SlIDA |
acts to induce |
tapetum and septum PCD |
Solanum lycopersicum |
| functional collaborations among four layers in lipid synthesis and transport |
are largely unknown |
anther development |
|
| tapetum |
contributes to |
pollen wall formation |
|
| anther En chloroplasts |
demonstrate essential roles in |
anther development |
Zea mays |
| Nin88 |
is expressed in |
tapetum layer |
Nicotiana tabacum |
| Arabidopsis (ATGPAT1, GPAT1, sn-2-GPAT1, AT1G06520) |
is important for |
nutrient secretion |
Arabidopsis thaliana |
| temporary survival of mutant anther |
and elongation stops |
mutant anther |
Zea mays |
| knowledge of production, transport, distribution, and utilization of energy and substances in four anther wall layers |
remains limited |
anther development |
|
| anther En chloroplasts |
function as sink organs for starch storage at early stages (7–9) and as photosynthetic factories for carbon fixation since stage 10 |
anther development |
Zea mays |
| OsMED14_1 transcripts |
are present in |
microspores and tapetum of early developing anthers |
Oryza sativa |
| WT tapetum thickness from stages 8b–11 |
gradually decreased |
stages 8b–11 |
Zea mays |
| defects of lipid metabolism in ER of tapetum |
can interrupt |
normal anther growth |
Zea mays |
| OsMED14_1 transcripts |
are not present in |
mature anther |
Oryza sativa |
| proper timing of ROS signaling in the septum and endothecium |
is required for |
pollen development and release |
|
| GhFLA19-D |
might be |
pivotal gene for normal development of anthers |
Gossypium hirsutum |
| SlIDA promoter-β-glucuronidase (GUS) fusion construct |
shows GUS activity that decreases at |
dehiscence stage |
Solanum lycopersicum |
| SlIDA |
may have a function during |
anther development |
Solanum lycopersicum |
| retarded septum PCD |
leads to |
dehiscence defect |
Solanum lycopersicum |
| three genes (GH_D12G1039, GH_D12G1042, GH_D12G1051) |
were found to be preferentially expressed in |
anthers |
Gossypium hirsutum |
| anther wall |
contains |
endothecium (En) |
|
| WT tapetum thickness from stages 7–8b |
gradually increased |
stages 7–8b |
Zea mays |
| CR-slida tapetum |
showed significant expansion at |
microspore stage |
Solanum lycopersicum |
| SlIDA |
is expressed in |
pollen |
Solanum lycopersicum |
| ectopic application of the SlIDA peptide |
rescued |
CR-slida anther dehiscence and PCD phenotypes |
Solanum lycopersicum |
| GbFLA19s in G. barbadense Hai 7124, Hai 1, and 3-79 |
were expressed at |
relatively equilibrated levels |
Gossypium barbadense |
| (ATMYB35, MYB35, TDF1, AT3G28470) (AMS, AT2G16910) and (AtMYB103, ATMYB80, MS188, MYB103, MYB80, AT5G56110) |
are |
key upstream transcription factors (TFs) for regulation of tapetal development and exine formation |
Arabidopsis thaliana |
| lack of effects on the TFs (ATMYB35, MYB35, TDF1, AT3G28470) (AMS, AT2G16910) and (AtMYB103, ATMYB80, MS188, MYB103, MYB80, AT5G56110) |
indicated that |
GoFLA19s may be downstream regulators of these TFs or modifiers in other pathways |
Gossypium hirsutum |
| CSA (Carbon Starved Anther) |
is |
key regulator of anther development |
|
| ChIP-Seq technology and bioinformatics |
identified |
downstream genes of transcription factors |
|
| normal structure and function of endothecium (En) chloroplasts maintained by ZmMs33-mediated lipid biosynthesis in tapetal cells |
are crucial for |
maize anther development |
Zea mays |
| outgrowths that ultimately form the four locules |
develop from cells located in between |
adaxial and abaxial domains |
Oryza sativa |
| cell layers |
could sometimes be recognized in |
(ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– anthers |
|
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– mutant tapetum |
did not form a thin and darkly stained ring around the microspores, but remained thick and disorganized |
tapetal cell organization |
|
| (AMS, AT2G16910) |
is important for |
anther development |
Arabidopsis thaliana |
| ER/ (ERL1, AT5G62230) (ERL2, AT5G07180) genes |
function in promoting |
longitudinal cell divisions |
|
| male sterility in ccc mutant |
is due to |
lack of lignification in anther endothecium |
Arabidopsis thaliana |
| ZmMs33 / ZmGPAT6 |
is expressed mainly in |
tapetum at early anther developmental stages |
Zea mays |
| chloroplasts in En cells of plant anthers |
have been reported in |
maize |
Zea mays |
| CR-slida lines |
show little effect on |
secondary wall lignification in the anther endothecium |
Solanum lycopersicum |
| SlIDA function in altering ROS levels |
might be limited to |
septum cell |
Solanum lycopersicum |
| GoFLA19s |
contribute to |
tapetal development |
Gossypium spp. |
| expression signal of GhFLA19s |
obviously decreased and was restricted to |
callose layers surrounding tetrad |
Gossypium hirsutum |
| non-functional GoFLA19s in hybrid |
leads to |
male sterility |
Gossypium hirsutum; Gossypium barbadense |
| OsMTR1 |
is predominantly expressed in |
early developmental stage anthers |
Oryza sativa |
| parietal cells |
differentiate into |
inner anther walls |
|
| diverse expression changes in exine development-related genes |
imply |
complex regulatory pathway affected by GoFLA19s during male reproductive development in cotton |
Gossypium hirsutum |
| endothecium (EN) |
is essential for |
anther dehiscence and pollen release |
|
| Rice OsGPAT3 and maize ZmMs33/ZmGPAT6 |
are different from |
(ATGPAT3, GPAT3, AT4G01950) and (ATGPAT6, GPAT6, AT2G38110) in Arabidopsis |
Oryza sativa; Zea mays; Arabidopsis thaliana |
| RBSDV P7-1 overexpression |
caused |
non-dehiscent anthers |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) cascade |
has been reported to function in |
anther development |
Arabidopsis thaliana |
| GhFLA19-D |
showed significantly higher expression levels than |
GhFLA19-A |
Gossypium hirsutum |
| OsMTR1 expression pattern |
implies |
diverse roles of FLA19s in dicots |
|
| GhFLA19-D |
showed |
persistent high expression levels in all stages up to mature stage |
Gossypium hirsutum |
| irregular expression of sporopollenin synthesis genes |
indicates |
GoFLA19s affect pathways that regulate expression of genes required for tapetal development and pollen exine formation |
Gossypium hirsutum |
| GoFLA19s |
play |
crucial roles in tapetal differentiation and degeneration |
Gossypium hirsutum |
| GhFLA19-D and GhFLA19-A |
showed significantly higher expression levels in |
G. hirsutum CCRI040029 and TM-1 |
Gossypium hirsutum |
| ms33-6038 mutant |
lacks |
exserted anthers at anthesis |
Zea mays |
| tapetum |
involved in |
subcellular organelle development |
|
| mpk6-1 anther |
produced less pollen and was smaller in size than |
(ATMAPK3, ATMPK3, MPK3, AT3G45640) mutant and wild-type anthers |
|
| abnormal anther phenotypes |
include |
abnormal anther cell differentiation |
Arabidopsis thaliana |
| slightly different growth conditions |
can impact |
normal anther development |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– mutant |
displays reduced expression of |
(AMS, AT2G16910) |
Arabidopsis thaliana |
| er-105 erl1-2 erl2-1 triple mutant |
produced stamens with |
apparently undifferentiated anthers of variable sizes |
Arabidopsis thaliana |
| er-105 erl1-2 erl2-1 triple mutant |
is defective in |
anther differentiation |
Arabidopsis thaliana |
| RNase activities in 30-40 kDa range in petunia anthers |
are only observed in |
early stages of anther development |
Petunia hybrida |
| (NZZ, SPL, AT4G27330) mutant |
fails to form |
L2 derived somatic cell types |
Arabidopsis thaliana |
| related genes ( (ATMYB35, MYB35, TDF1, AT3G28470) (AMS, AT2G16910) (AtMYB103, ATMYB80, MS188, MYB103, MYB80, AT5G56110) (DRL1, TKPR1, AT4G35420) (ATLAP3, LAP3, AT3G59530) ) |
are still expressed regularly in |
mutant (ghfla19-1) |
Gossypium hirsutum |
| 24-nt phasiRNAs |
accumulate at later anther developmental stages compared to |
21-nt phasiRNAs |
|
| (ATMYB33, MYB33, AT5G06100) (ATMYB65, MYB65, AT3G11440) mutants |
tapetum becomes highly vacuolated in at |
anther stage 5 |
Arabidopsis thaliana |
| er-105 erl1-2 erl2-1 mutant |
resembles |
known loss-of-function Arabidopsis tapetum mutants |
Arabidopsis thaliana |
| residual function of ER/ERL and (ATMAPK3, ATMPK3, MPK3, AT3G45640) /6 genes |
may be near |
threshold of needed activity for archesporial formation or function |
Arabidopsis thaliana |
| adaxial lobes |
develop slightly later than |
abaxial lobes |
Arabidopsis thaliana |
| Hsp17-CII transcripts |
are strongly accumulated in |
anthers at young stages |
Solanum lycopersicum |
| lack of lignification in anther endothecium |
causes |
failure of anther dehiscence |
Arabidopsis thaliana |
| C3'H co-suppressed Arabidopsis lines |
exhibit |
male sterility |
Arabidopsis thaliana |
| four layers of anther wall |
are finely regulated throughout |
anther development |
|
| epidermis |
is located at |
outermost layer of the anther |
|
| autophagy |
is crucial for |
anther dehiscence and male fertility in plant anthers |
|
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) /– mutant |
characterized for |
anther development |
Arabidopsis thaliana |
| er-105 erl1-2 erl2-1 triple mutant |
exhibits |
more severe anther phenotypes |
Arabidopsis thaliana |
| (EMS1, EXS, AT5G07280) |
expression not significantly different in |
(ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– mutant flowers |
Arabidopsis thaliana |
| erl1-2 single mutant |
had anthers that were comparable in size to |
wild-type anthers |
Arabidopsis thaliana |
| er-105 erl1-2 erl2-1 triple mutant anthers that produced viable pollen |
were larger than |
anthers that did not differentiate |
Arabidopsis thaliana |
| er-105 erl1-2 erl2-1 triple mutant anthers |
had phenotypes varying from |
antherless stamens to anthers with no more than two abaxial lobes with differentiated cell layers |
Arabidopsis thaliana |
| er-105 erl1-2 erl2-1 mutant |
lack of ER/ERL1/ERL2 function leads to |
aberrant cell patterning |
|
| high temperature at anthesis |
affects |
anther protein expression |
Oryza sativa L. |
| anther endothecium |
undergoes expansion |
expansion |
|
| abnormal tapetum behaviour in Allium |
was characterized by |
extremely early degeneration and tapetum hypertrophy and autolysation |
Allium |
| outgrowths that ultimately form the four locules |
are associated with |
boundaries |
Oryza sativa |
| er-105 erl1-2 erl2-1 anthers |
were not observed to |
dehisce |
Arabidopsis thaliana |
| ER/ (ERL1, AT5G62230) (ERL2, AT5G07180) |
play important roles in |
anther lobe formation |
Arabidopsis thaliana |
| under-developed anthers of er-105 erl1-2 erl2-1 triple mutant |
were overall |
oval or kidney-shaped |
Arabidopsis thaliana |
| (EMS1, EXS, AT5G07280) and (TPD1, AT4G24972) expression |
was not reduced in |
(ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) /– mutant |
Arabidopsis thaliana |
| ER/ (ERL1, AT5G62230) (ERL2, AT5G07180) |
play important roles in |
anther cell differentiation |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) |
expression level not significantly different in |
(EMS1, EXS, AT5G07280) mutant |
Arabidopsis thaliana |
| mutant middle-layer cells at stage 6 in er-105 erl1-2 erl2-1 triple mutant |
were larger than in wild-type, suggesting |
they persisted longer than normal |
Arabidopsis thaliana |
| ER/ (ERL1, AT5G62230) (ERL2, AT5G07180) genes |
contribute to |
normal cell patterning |
|
| OsPBP1 antisense transgenic line A-38 |
had |
shrunken, short, and bended anthers |
Oryza sativa |
| ER/ (ERL1, AT5G62230) (ERL2, AT5G07180) and (ATMAPK3, ATMPK3, MPK3, AT3G45640) (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) |
might function in the same pathway to regulate |
anther cell division and differentiation |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) /– mutant |
had later defects in |
differentiation of anther cell layers |
|
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /6-dependent signal transduction |
promote |
archesporial formation or function |
Arabidopsis thaliana |
| (ATHSFA2, HSFA2, AT2G26150) |
transcript levels are more detectable during development at |
2 mm anther stage under control conditions |
Solanum lycopersicum |
| Solyc08g062780.1.1 (AtAMS-like and OsTDR-like) |
is probably involved in |
tapetum degeneration |
Solanum lycopersicum |
| Arabidopsis bHLH89/91 |
are suggested to be involved in |
tapetal PCD |
Arabidopsis thaliana |
| RNA-seq data |
should provide good basis for |
identification and analysis of new genes involved in anther development in tomato |
Solanum lycopersicum |
| decrease in (ATMYB33, MYB33, AT5G06100) gene expression |
correlated with increase in miR159 gene expression would cause |
anther defects, male sterility, and delayed flowering |
|
| strong expression signal of GhFLA19s |
was detected predominantly in |
tapetum and microspores during meiosis stage |
Gossypium hirsutum |
| anther |
contains |
tapetum and microsporocytes |
|
| five cell layers within each lobe |
are, from outer to inner |
epidermis, endothecium, middle layer, tapetum, and pollen mother cells (PMCs) |
Arabidopsis thaliana |
| clusters of cells at the four corners |
begin to establish |
cell layers parallel to the epidermis |
|
| locule of er-105 erl1-2 erl2-1 triple mutant anther |
had |
five cell layers |
Arabidopsis thaliana |
| ER/ (ERL1, AT5G62230) (ERL2, AT5G07180) genes |
function together to regulate |
anther lobe formation |
|
| ER, (ERL1, AT5G62230) and (ERL2, AT5G07180) |
redundantly promote |
propagation of cells necessary for normal anther formation |
|
| (ATHSFA2, HSFA2, AT2G26150) |
was expressed early in |
young anthers |
Solanum lycopersicum |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) /– mutant |
exhibits |
less severe anther phenotypes |
Arabidopsis thaliana |
| tapetum and middle-layer cells |
may be delayed in |
their development |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– anther |
formed |
all four lobes |
|
| single, double and er-105 erl1-2 (+/–) erl2-1 mutants |
show |
normal anther development |
|
| failure to form an anther lobe |
could be due to |
absence or abnormal function of archesporial cells |
Arabidopsis thaliana |
| microsporogenesis in petal-like organs of Arabidopsis agamous mutant |
often produces |
two locules |
Arabidopsis thaliana |
| Arabidopsis Tapetal Development and Function1 (ATMYB35, MYB35, TDF1, AT3G28470) |
is highly expressed in |
meiocytes |
Arabidopsis thaliana |
| temperature |
does not affect |
anther size |
Oryza sativa |
| Hsp17.4-CII |
transcript levels are more detectable during development at |
2 mm anther stage under control conditions |
Solanum lycopersicum |
| Hsp17.6-CII |
is expressed particularly in |
2 mm anthers |
Solanum lycopersicum |
| (TPD1, AT4G24972) |
is involved in |
tapetum development |
Arabidopsis thaliana |
| ER/ (ERL1, AT5G62230) (ERL2, AT5G07180) genes |
may communicate signals necessary for |
normal tapetum cell division |
|
| anther wax layer |
protecting anther from |
various environmental stresses |
Oryza sativa |
| (AMS, AT2G16910) |
is strongly expressed in |
tapetum cells specifically after meiosis |
Arabidopsis thaliana |
| ms10 35 mutant |
showed |
protruded stigma |
Solanum lycopersicum |
| growth of tapetal cells |
was more advanced than |
control |
Lilium longiflorum |
| er-105 erl1-2 erl2-1 triple mutant |
appears to have |
more severe phenotype than the (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) /– mutant |
Arabidopsis thaliana |
| single, double and er-105 erl1-2 (+/–) erl2-1 mutants |
produced |
five anther cell layers |
Arabidopsis thaliana |
| degradation of some of tapetal cells in er-105 erl1-2 erl2-1 triple mutant |
possibly due to |
tapetum did not completely surround meiocytes |
Arabidopsis thaliana |
| ER, (ERL1, AT5G62230) and (ERL2, AT5G07180) |
have overlapping expression during |
anther development |
|
| er-105 erl1-2 erl2-1 mutant |
lack of ER/ERL1/ERL2 function leads to |
increased numbers of tapetum cells |
|
| archesporial cells |
are formed in |
L2 layer of the anther primordia |
Arabidopsis thaliana |
| genes that directly control anther development, such as (NZZ, SPL, AT4G27330) |
are vulnerable to factors that disrupt normal activity of |
these genes in the absence of ER/ERL and (ATMAPK3, ATMPK3, MPK3, AT3G45640) /6 gene functions |
Arabidopsis thaliana |
| (ATHSFA2, HSFA2, AT2G26150) and Hsp17-CII |
are finely regulated during |
anther development |
Solanum lycopersicum |
| heat stress |
causes failure of |
adequate dehiscence in tomato anthers |
Solanum lycopersicum |
| callose |
regulates |
optimal extracellular distance and cell connectivity among central anther locular cells |
Oryza sativa |
| erl1-2 erl2-1 double mutant |
had anthers that were comparable in size to |
wild-type anthers |
Arabidopsis thaliana |
| ER/ERL and (ATMAPK3, ATMPK3, MPK3, AT3G45640) /6 genes |
may increase |
robustness of regulatory machinery of early anther development |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– mutant anther |
was even smaller than |
either single mutant anther |
|
| wild-type anthers |
are roughly oval-shaped at |
anther stage 2 |
|
| er-105 erl1-2 erl2-1 triple mutant |
produced fewer |
stamens |
Arabidopsis thaliana |
| (EMS1, EXS, AT5G07280) (ATSERK1, SERK1, AT1G71830) (ATSERK2, SERK2, AT1G34210) and (TPD1, AT4G24972) genes |
likely function in the same pathway to specify |
tapetum formation |
Arabidopsis thaliana |
| tapetum |
neighbors |
microsporocytes |
|
| Btr1-like |
is exclusively expressed in |
developing anthers |
Hordeum vulgare |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) /– mutant anthers |
sometimes do not |
dehisce |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) /– mutants |
have anthers that do not form |
several normal cell types |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– mutant |
produced anthers that were smaller than |
wild-type anthers |
|
| (ATMYB33, MYB33, AT5G06100) |
expression not significantly different in |
(ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– mutant flowers |
Arabidopsis thaliana |
| heat stress |
causes alterations in |
tomato anthers |
Solanum lycopersicum |
| SPATULA (SPT, AT4G36930) |
is strongly expressed in |
developing anthers |
Arabidopsis thaliana |
| (ATSERK1, SERK1, AT1G71830) and (ATSERK2, SERK2, AT1G34210) |
are expressed during |
anther development |
Arabidopsis thaliana |
| control anthers at young microspore stage |
show partial tapetal cell degeneration and tapetum becoming |
thin |
Nicotiana tabacum |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) and the ER-family genes |
are important for |
cell differentiation during anther development |
Arabidopsis thaliana |
| middle-layer cells in er-105 erl1-2 erl2-1 triple mutant |
appear to overlap with one another |
triple mutant anther |
Arabidopsis thaliana |
| ER/ (ERL1, AT5G62230) (ERL2, AT5G07180) and (ATMAPK3, ATMPK3, MPK3, AT3G45640) (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) |
act in |
the same pathway important for normal anther development |
Arabidopsis thaliana |
| ER/ERL and (ATMAPK3, ATMPK3, MPK3, AT3G45640) /6 genes |
act to modulate |
intrinsic anther developmental program |
Arabidopsis thaliana |
| (EMS1, EXS, AT5G07280) |
is important for |
anther development |
Arabidopsis thaliana |
| symmetric cell divisions |
do not typically occur in the same plane as |
asymmetric cell division events |
|
| (NZZ, SPL, AT4G27330) and (BAM1, AT5G65700) (BAM2, BMY9, AT4G00490) |
are important for controlling |
cell differentiation of archesporial cell progeny |
Arabidopsis thaliana |
| hampered response to environmental changes |
results in |
developmental defects |
Arabidopsis thaliana |
| genes essential for anther development |
selected for |
expression analysis by real-time PCR |
|
| connective cell walls |
gradually degenerate during transition from mature to later phase |
female stage |
|
| GoFLA19s |
have specific expression in |
anthers |
Gossypium hirsutum |
| er-105 single mutant |
had anthers that were comparable in size to |
wild-type anthers |
Arabidopsis thaliana |
| er-105 erl1-2 erl2-1 triple mutant |
produced |
filament-like stamens that appeared to completely lack an anther |
Arabidopsis thaliana |
| tapetal cells in er-105 erl1-2 erl2-1 triple mutant when PMCs had developed into microspores |
remained predominately attached to each other and to other somatic cells |
stage 8 |
Arabidopsis thaliana |
| IR64 |
has |
longest and widest anthers with cross-sectional area of 2.44 mm² |
Oryza sativa |
| anthers from buds less than 1 mm in length |
contain |
sporophytic tissues |
|
| middle layers |
have degenerated during development |
female stage |
|
| GhFLA19-D |
was downregulated in |
CCRI9106 during anther development |
Gossypium hirsutum |
| signaling molecules |
play crucial roles in determining |
anther cell types |
Arabidopsis thaliana |
| (AtPAO5, PAO5, AT4G29720) |
is expressed in |
anther tapetal cells |
Arabidopsis thaliana |
| CsGAMYB1 overexpression in Arabidopsis wild-type Columbia |
results in |
smaller anthers |
Arabidopsis thaliana |
| tapetal cells in er-105 erl1-2 erl2-1 triple mutant |
were greater in number compared to |
wild-type |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) |
is not sufficient to consistently direct |
normal cell differentiation |
|
| er-105 erl1-2 erl2-1 and (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) /– anthers |
can still form |
tapetum |
Arabidopsis thaliana |
| PMCs |
degenerate |
in (ATMYB33, MYB33, AT5G06100) (ATMYB65, MYB65, AT3G11440) and (DYT1, AT4G21330) mutants |
Arabidopsis thaliana |
| er (ERL1, AT5G62230) (ERL2, AT5G07180) mutants |
lack |
consistent anther morphologies |
Arabidopsis thaliana |
| SPATULA (SPT, AT4G36930) |
is expressed in |
tapetum |
Arabidopsis thaliana |
| mutant anthers |
produced |
four lobes |
|
| asymmetric cell divisions |
occur |
anther locule formation |
|
| ER/ERL and (ATMAPK3, ATMPK3, MPK3, AT3G45640) /6 |
may modulate |
(NZZ, SPL, AT4G27330) expression |
Arabidopsis thaliana |
| ER/ERL and (ATMAPK3, ATMPK3, MPK3, AT3G45640) /6 |
are not required for |
(NZZ, SPL, AT4G27330) expression |
Arabidopsis thaliana |
| dirigent-like protein |
could influence |
anther dehiscence |
Oryza sativa |
| Hsp17.4-CII |
is expressed during |
anther development |
Solanum lycopersicum |
| understanding of anther development processes |
may provide |
effective mechanisms for efficient generation of hybrids |
|
| control anthers at young pollen to pollen mitosis stage |
show almost all tapetal cells differentiated and |
degenerated |
Nicotiana tabacum |
| proteases |
play an important role in |
tapetum degradation |
|
| connective cell wall |
is well developed at |
immature stage |
|
| multinucleated periplasmodium |
degenerates and finally disappears |
during anther development |
|
| such layers |
were not well organized or were otherwise abnormal |
cell layer organization |
|
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– mutant |
play crucial roles in |
anther cell differentiation |
|
| ER/ (ERL1, AT5G62230) (ERL2, AT5G07180) and (ATMAPK3, ATMPK3, MPK3, AT3G45640) /6 genes |
define |
new function in regulation of anther development |
Arabidopsis thaliana |
| ER/ERL and MPK genes |
likely mediate |
cell–cell communication |
Arabidopsis thaliana |
| thickening in the endothecium |
limits expansion |
expansion |
|
| male sterility |
is due to |
defects in anther dehiscence |
Arabidopsis thaliana |
| anti-NtCP56 transgenic anthers from young pollen stage |
show endothecium cells remaining loosely arrayed and lacking |
clear fibrotic cell walls |
Nicotiana tabacum |
| mutations in (EMS1, EXS, AT5G07280) |
cause formation of |
extra microsporocytes |
Arabidopsis thaliana |
| tapetum of MYB35-SRDX anthers at stage 8 |
degenerated earlier than |
wild-type tapetum |
Arabidopsis thaliana |
| anther dehiscence |
involves |
changes to the water status of the anther |
|
| tapetum |
is visible at |
immature stage |
|
| tapetal cells in Arabidopsis mutant (MS1, AT5G22260) |
became |
highly vacuolated in the early stages |
Arabidopsis thaliana |
| microspore release |
occurs before |
anther endothecium expansion and deposition of ligno-cellulosic secondary thickening |
|
| typical mitochondria |
were not observed in |
tapetal cells of sterile anthers |
Brassica napus |
| extensive anther development |
occurs during |
inflorescence development |
|
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /– mpk6-1/+ anthers |
appeared to develop |
normal cell layers, similar to wild-type |
|
| (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) |
expressed at slightly higher level than |
(ATMAPK3, ATMPK3, MPK3, AT3G45640) |
Arabidopsis thaliana |
| er-105 erl1-2 erl2-1 triple mutant |
is defective in |
anther formation |
Arabidopsis thaliana |
| tapetal cells in er-105 erl1-2 erl2-1 triple mutant |
did not form |
single layer |
Arabidopsis thaliana |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ (ATMAPK6, ATMPK6, MAPK6, MPK6, AT2G43790) /– mutant tapetum |
had |
enlarged cells |
|
| establishment of endothecium thickenings |
was |
normal in knockout and knockdown lines |
Arabidopsis thaliana |
| breakdown of connective cell wall |
is completed and pollen grains are released at |
male stage |
|
| Solyc02g079810 |
is expressed only in |
anthers |
Solanum lycopersicum |
| Arabidopsis bHLH89/91 |
interact with |
(AMS, AT2G16910) |
Arabidopsis thaliana |
| secondary thickening of cell walls in endothecium |
is similar in |
wild type, AIF-C+SRDX, and AIF-C anthers at stage 11 |
Arabidopsis thaliana |
| 32kDa protein |
recognized only at |
microspore stage during anther development |
Lilium longiflorum |
| anther cells |
initiate specification and differentiation to form |
bilaterally symmetrical structure with four lobes |
Arabidopsis thaliana |
| Ms10 35 |
is expressed exclusively in |
meiocyte and tapetal tissues |
Solanum lycopersicum |
| sporogenous cell |
is |
innermost cell layer |
Arabidopsis thaliana |
| Ms10 35 and its homologues |
have conserved role in |
tapetum development |
|
| (ACO4, EAT1, EFE, AT1G05010) |
interacts with |
(GAMMA-TIP2, SITIP, TIP1;2, TIP2, AT3G26520) and (AtERF98, AtTDR1, ERF98, TDR1, AT3G23230) |
Oryza sativa |
| function of bHLH89/91 |
is not clear due to |
lack of genetic and molecular evidence |
Arabidopsis thaliana |
| Solyc03g113530.2.1 (AtTDF-like) |
was significantly downregulated in |
ms10 35 mutant |
Solanum lycopersicum |
| anther dehiscence |
is |
multistage process involving localized cellular differentiation and degeneration |
|
| (PXY, TDR, AT5G61480) mutant rice |
shows |
middle layer cell persistence |
Oryza sativa |
| GFP fluorescence |
becomes visible just after |
appearance of locules in anthers |
Arabidopsis thaliana |
| rice (GAMMA-TIP2, SITIP, TIP1;2, TIP2, AT3G26520) |
regulates |
rice (AtERF98, AtTDR1, ERF98, TDR1, AT3G23230) and (ACO4, EAT1, EFE, AT1G05010) |
Oryza sativa |
| OsGAMYB |
is involved in |
exine formation |
Oryza sativa |
| LLA1271 spatial and temporal expression patterns |
are well correlated with |
tapetal development and degeneration |
Lilium longiflorum |
| lily genes |
are co-expressed in |
tapetum and microspores |
Lilium longiflorum |
| Arabidopsis male sterile mutants |
analysis of enables understanding of |
gene regulatory networks controlling maternal development of anther |
Arabidopsis thaliana |
| archesporial cells |
divide periclinally to give rise to |
sporogenous cells |
|
| multinucleated periplasmodium cells |
contain |
dense cytoplasm with abundant mitochondria and plastids |
|
| cells of the triple mutant anthers |
appeared larger and more disorganized than |
wild-type |
Arabidopsis thaliana |
| archesporial cells |
divide to form |
the four anther lobes |
Arabidopsis thaliana |
| tapetal cells |
were disorganized with |
abnormally large vacuoles |
Arabidopsis thaliana |
| anther epidermis |
is covered with |
layer of wax |
Oryza sativa |
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– mutant anthers |
are defective in |
cell differentiation |
|
| each of the four lobes |
has |
four distinct, concentric layers of cells surrounding the pollen mother cells |
|
| (ATMAPK3, ATMPK3, MPK3, AT3G45640) /+ mpk6-1/– anther locules |
occasionally displayed |
similar cell patterning and structure to wild-type |
|
| (NZZ, SPL, AT4G27330) |
is important for |
anther development |
Arabidopsis thaliana |
| tapetal cells in er-105 erl1-2 erl2-1 triple mutant |
were greatly enlarged and abnormally shaped |
triple mutant anther |
Arabidopsis thaliana |
| er (ERL1, AT5G62230) (ERL2, AT5G07180) triple mutant |
produced |
reduced number of locules |
Arabidopsis thaliana |
| anther primordial cells |
generate |
meiocytes and four-lobed cell layers |
Oryza sativa |
| (ATHSFA2, HSFA2, AT2G26150) |
expression in 2 mm anthers from control plants is higher than in |
older anthers |
Solanum lycopersicum |
| mutations in (TPD1, AT4G24972) |
cause failure to form |
tapetal cells |
Arabidopsis thaliana |
| tapetum function |
is tightly coupled with |
progress of meiosis and pollen maturation |
|
| (AMS, AT2G16910) |
is |
essential for anther development |
|
| Ogura cytoplasmic male sterility (CMS) |
shows |
decrease in tapetal cell stainability |
Raphanus sativus |
| tapetum in arum species |
is known to be |
amoeboid |
|
| enzymatic lysis of the septum |
occurs before |
programmed cell death-like breakdown of the septum |
|
| Tapetal mitochondrial alterations |
were also reported in |
male-sterile anthers of Petunia hybrida cv. Blue Bedder |
Petunia hybrida |
| ERECTA |
is involved in |
anther development |
Arabidopsis thaliana |
| new role for (ACBP3, AT4G24230) in the development of anthers in Arabidopsis Ler-0 |
is reported |
anther development in Arabidopsis Ler-0 |
Arabidopsis thaliana |
| HvTDF1 |
expression starts from |
anther stage 7 |
Hordeum vulgare |
| (AtC3H15, CDM1, AT1G68200) mutation |
did not affect |
tapetum morphology |
Arabidopsis thaliana |
| ACBP3pro (Col)::ACBP3::GFP |
did not show high expression in |
anthers |
Arabidopsis thaliana |
| knockout mutants of genes affecting (NZZ, SPL, AT4G27330) expression or those regulated by such as BRASSINAZOLE-RESISTANT (BZR), TRYPTOPHAN AMINOTRANSFERASE OF ARABIDOPSIS 1 (CKRC1, SAV3, TAA1, TIR2, WEI8, AT1G70560) TRYPTOPHAN AMINOTRANSFERASE RELATED2 (AtTAR2, TAR2, AT4G24670) and (bZIP21, TGA9, AT1G08320) (bZIP65, TGA10, AT5G06839) |
exhibit similar phenotypes as |
(NZZ, SPL, AT4G27330) mutant |
Arabidopsis thaliana |
| (AtPAO5, PAO5, AT4G29720) |
is expressed in |
anther-filament junction |
Arabidopsis thaliana |
| (AtERF98, AtTDR1, ERF98, TDR1, AT3G23230) mutant |
exhibits |
male sterility |
Oryza sativa |
| OsTDR1 |
is regulated by |
OsUDT1 |
Oryza sativa |
| gibberellin (GA) |
up-regulates |
LLA1271 gene |
Lilium longiflorum |
| five (bHLH, AT5G51780) genes ( (DYT1, AT4G21330) (AMS, AT2G16910) (bHLH089, AT1G06170) (bHLH091, AT2G31210) and (bHLH010, AT2G31220) ) |
were found to be preferentially expressed in |
anther |
Arabidopsis thaliana |
| restored plants |
showed |
more vacuolization of the tapetum |
Brassica napus |
| Capsicum annuum L. cytoplasmic male-sterile line CMS 21A |
showed |
abnormally swollen tapetum pressed against the uni-nucleate microspores |
Capsicum annuum |
| NtCP56 |
might be involved in |
anther development |
Nicotiana tabacum |
| HvTDF1 |
has |
tapetum-specific expression during anther stage 7/8 |
Hordeum vulgare L. |
| (ACBP2, AtACBP2, AT4G27780) |
has precise role in anther development that remains to be elucidated |
anther development |
Arabidopsis thaliana |
| difference in expression alteration of β-1,3-glucanase genes at different anther stages |
suggested that they potentially function at |
different anther stages |
|
| ERECTA |
modulates |
(ACBP3, AT4G24230) function |
Arabidopsis thaliana |
| male sterile mutants in maize |
isolated in |
male sterility screens |
Zea mays |
| Ms32 |
inhibits |
further periclinal division in differentiated layers |
|
| two amino acids inserted in the ms32-6066 mutant protein |
may disrupt |
binding to DNA |
Zea mays |
| GFP signal |
is stronger in |
tapetal cells at stage 6 |
|
| (DYT1, AT4G21330) expression |
occurs at similar tapetal development stages as |
(ATMYB35, MYB35, TDF1, AT3G28470) expression |
|
| post-meiotic haploid male gametophytes |
have been evaluated |
by transcriptome analysis |
Oryza sativa; Zea mays; Arabidopsis thaliana |
| Male sterile 1035 (Ms1035) |
regulates genes involved in |
transport |
Solanum lycopersicum |
| (ATMYB26, MS35, MYB26, AT3G13890) |
have been implicated in |
secondary thickening of anther endothecium and anther dehiscence |
|
| pollen maturation |
begins when |
epidermis, endothecium, middle layer and tapetum inside the anther have differentiated, and meiosis has been completed |
Arabidopsis thaliana |
| endothecium lignification |
is controlled by |
auxin |
|
| IAA concentration |
increases |
meiotic and pre-meiotic stages |
Arabidopsis thaliana |
| presumptive middle layer in ms32-ref |
is affected by |
absence of MS32 |
|
| ms32-6066 allele |
has |
two amino acid insertion in the (bHLH, AT5G51780) domain |
Zea mays |
| down-regulation of most DNA replication genes |
may contribute to |
the delay in cell divisions observed in some cell types |
Zea mays |
| msca1 mutant anthers |
lack differentiation of |
somatic L2-d anther cell types |
Zea mays |
| (AtMYB103, ATMYB80, MS188, MYB103, MYB80, AT5G56110) |
has important role in |
callose dissolution |
Arabidopsis thaliana |
| anther indehiscence |
occurs during |
high-temperature (HT) stress |
|
| (AP2, AtAP2, FL1, FLO2, AT4G36920) |
was suppressed at |
TS but induced at the TDS in H05 |
Gossypium hirsutum |
| wild-type samples |
were collected from |
five distinct anther stages, stage 6–8b |
Hordeum vulgare |
| hvtdf1 stage 8a2 and 8b |
had a closer correlation with |
the WT stage 7 data rather than the later stages |
Hordeum vulgare |
| acbp3-2 mutant |
exhibits |
defective adaxial anther lobes |
Arabidopsis thaliana |
| defects in Landsberg anther development |
related to |
ERECTA-mediated pathway |
Arabidopsis thaliana |
| Osalkbh5 anthers at Stage 10 |
show |
thick tapetal layers |
Oryza sativa L. ssp. japonica |
| HvTDF1 gene expression |
was detected in the tapetum from |
Microspore Mother Cell (MMC) stage and tetrad |
Hordeum vulgare |
| dyads formed in the mutant line at early single microspore stage |
became vacuolated during the subsequent development stages until |
the anther locule collapsed |
Hordeum vulgare |
| For barley, two comparison groups, hvtdf1 stage 8a2 vs WT stage 7, and hvtdf1 stage 8b vs WT stage 8a1 |
were selected based on |
the delayed development between WT and mutant plants |
Hordeum vulgare |
| Ms10 35 may affect stamen length |
by regulating |
Se2.1 |
Solanum lycopersicum |
| AIF-C+VP16 plants |
show promoted |
anther dehiscence |
Arabidopsis thaliana |
| uniconazole and NBD treatment |
may affect |
anther gene expression |
Lilium longiflorum |
| endothecium |
is |
second cell layer from outer |
Arabidopsis thaliana |
| weak band of 22kDa protein |
detected at |
pre-meiotic phase of anther development |
Lilium longiflorum |
| LLA1271 gene |
is negatively regulated by |
ethylene |
Lilium longiflorum |
| protruded stigma in ms10 35 |
was due to |
significantly reduced anther cone size |
Solanum lycopersicum |
| (bHLH, AT5G51780) subfamilies containing Ms10 35 and AtAMS |
have conserved function |
anther development |
|